Biennials or perennials; (caudex branched); usually densely hirsute (at least basally), trichomes simple or stalked, forked, sometimes glabrescent with mostly simple ones. Stems simple or several from base (rosette), erect, often branched distally, 2-7(-10) dm, (hirsute basally, trichomes simple and minutely stalked, forked, rarely almost exclusively glabrous). Basal leaves: petiole 0.7-3 cm, (ciliate or not); blade broadly spatulate to narrowly oblanceolate, 1.2-10 cm × 7-30 mm, margins entire or dentate, apex obtuse or acute, surfaces usually pubescent, sometimes glabrous, trichomes simple, forked and stalked, stellate. Cauline leaves (2-)4-12 (-18), (well-spaced or, rarely, overlapping); blade ovate to oblong or lanceolate, 1.5-6.5(-10) cm × 5-30 mm, base auriculate, margins dentate or entire, apex acute or obtuse, surfaces sparsely hirsute or glabrous. Racemes simple or branched. Fruiting pedicels erect to erect-ascending, 3.5-10(-15) mm, (glabrous or sparsely pubescent). Flowers: sepals oblong, 3.5-5 × 1.2-2 mm, lateral pair saccate basally; petals white, rarely pink, linear-oblanceolate or narrowly spatulate, (5.5-) 6.5-9(-10) × 1.5-3 mm, apex obtuse; filaments 3.5-6 mm; anthers oblong, 0.7-1 mm. Fruits erect to erect-ascending, (often appressed to rachis), torulose, 3.5-6.5 cm × 1.2-1.8(-2) mm; valves each with prominent midvein extending full length or to middle; ovules 54-80 per ovary; style (0.1-)0.3-1 mm. Seeds usually narrowly winged throughout or only distally, rarely not winged, oblong or suborbicular, 1-1.8 × 0.9-1.3 mm; wing to 0.2 mm wide distally. 2n = 32, 64.
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