Arachniodes simplicior (Makino) Ohwi

Simpler hollyfern (en)

Species

Pteridophytes > Polypodiales > Dryopteridaceae > Dryopteridoideae > Arachniodes

Characteristics

Rhizome shortly creeping, stiff, apex densely scaly; scales (reddish or dark) brown, linear-lanceolate or subulate, ca. 8 × 0.5-1.5 mm, base often remotely fimbriate, apex hairlike and often tortuous, firmly membranous. Fronds approximate, 0.4-1.1 cm; stipe stramineous, 20-65 cm, ca. 3 mm in diam., base with scales similar to those on rhizome, scales sparser upward, or rarely glabrescent; lamina (2 or)3-pinnate, pale green when dried, ovate-oblong, deltoid-ovate, or pentagonal, 20-60 × 15-40 cm, firmly papery or leathery, base rounded, cordate, or hastate, apex abruptly caudate with a stalked conform terminal pinna, rachis sparsely scaly as distal portion of stipe, or glabrescent throughout; lateral pinnae 2-5(or 6) pairs, alternate or lowest pair opposite, with to 2.5 cm stalk, obliquely spreading; lowest pinnae obliquely deltoid, to 28 × 15 cm, base broadly cuneate, apex caudate-acuminate; primary pinnules ca. 20 pairs, with to 1 cm stalk, basal pair elongated (basiscopic one longer, to 15 × 2.2 cm), lanceolate, base cuneate, apex acuminate, pinnate, larger fronds with proximal 2 pairs of pinnules elongated and pinnate; secondary pinnules shortly stalked or sessile, oblong or larger ones slightly falcate, ca. 1.5 × 0.6 cm, base acroscopically truncate and auriculate, basiscopically cuneate, apex acute, margin sharply serrate (or shallowly pinnatifid), aristate; upper 1-3 pairs of pinnae with basal pair of pinnules elongated, other lateral pinnae shorter and lanceolate; axes and veins of ultimate segments with minute (dark) brown subulate appressed scales abaxially. Sori terminal on veinlets, 1-6 pairs per ultimate segment, medial between midvein and margin or closer to margin; indusia (dark) brown, membranous, entire. 2n = 82*.
More
Stems creeping, 5--8 mm diam., densely scaly; scales tan to brownish, lanceolate. Leaves monomorphic, evergreen, 40--85 cm. Petiole straw-colored to tan, spaced 0.5--4 cm apart, 15--46 cm × 4--6 mm, base scaly; scales lanceolate. Blade deltate-pentagonal, 2--3-pinnate, 24--40 cm, broadest at base, abruptly tapering distally, apex pinnate, ultimately pinnatifid, as long as or longer than rest of blade. Pinnae 3--5 pairs, 1-pinnate, proximal pair basally 2-pinnate, 8--18 × 2--4 cm (excluding expanded base of proximal pinnae). Basal pinnules of proximal pair of pinnae elongate and pinnalike; basiscopic pinnule to 10 cm, acroscopic pinnule 1/3--1/2 length of basiscopic. Ultimate segments undivided to pinnatifid, finely spiny along margins and at tip. Sori with indusia 0.5--1 mm diam., indusia with deep sinus and often overlapping lobes, glabrous.
Life form perennial
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality -
Pollination -
Spread -
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.45 - 0.93
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light 1-6
Soil humidity -
Soil texture -
Soil acidity 2-8
Soil nutriment -
Hardiness (USDA) 9-11

Usage

Uses medicinal
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by divisions.
Mode divisions
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Arachniodes simplicior unspecified picture

Distribution

Arachniodes simplicior world distribution map, present in China, Japan, Korea (Democratic People's Republic of), United States of America, and Viet Nam

Identifiers

LSID urn:lsid:ipni.org:names:17427950-1
WFO ID wfo-0001110707
COL ID G33S
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Arachniodes simplicior Byrsopteris simplicior Rumohra simplicior Polystichopsis simplicior Arachniodes fujiangensis Arachniodes calcarata Polystichum simplicius Arachniodes yasu-inouei Arachniodes aristatissima Arachniodes parasimplicior Arachniodes tibetana Arachniodes jiulungshanensis Aspidium aristatum var. simplicior Polystichum aristatum var. simplicior