Arachniodes Blume

Hollyfern (en)

Genus

Pteridophytes > Polypodiales > Dryopteridaceae > Dryopteridoideae

Characteristics

Plants terrestrial, evergreen, medium-sized (or large). Rhizome long creeping to short and ascending, or rarely erect, densely scaly; scales filiform, subulate, lanceolate, or ovate, often attenuate toward apex, margin entire or with toothlike (rarely fimbriate) projections. Fronds few, remote or approximate; stipe equal in length to lamina or longer, base as densely scaly as rhizome, adaxially sulcate; lamina deltoid, ovate, pentagonal, or rarely lanceolate, usually 2-4(or 5)-pinnate, but simply pinnate in a few species, herbaceous, papery to leathery, glabrous or with a few narrow scales, rarely hairy; dissection anadromous throughout; rachis adaxially sulcate, grooves continuous with those on axes of higher order, scaly to glabrescent, or rarely with pale gray unicellular acicular hairs, scales entire (or with irregular projections on margin), sometimes with enlarged and denticulate base; pinnae a few to numerous pairs, usually shortly stalked, basal ones often with long basal basiscopic pinnules; upper divisions gradually reduced and ending in acuminate or caudate apices, or with a distinct terminal pinna; basiscopic pinnules often slightly longer than acroscopic ones; ultimate segments sessile, usually oblong to rhomboid, or rarely linear, often inequilateral, with an acroscopic auricle, margin sharply dentate to aristate; venation usually also anadromous, veins free, pinnate or forked, not quite reaching margin of ultimate segment, often ending in a hydathode. Sori orbicular, terminal or dorsal on acroscopic branches of veinlets; indusia orbicular-reniform, attached at a deep sinus, persistent or fugacious; sporangia numerous, long stalked, with an annulus of 13-16 thickened cells. Spores ellipsoid, often with broad rugate perispore. x = 41.
More
Plants terrestrial. Stems moderately long-to short-creeping, stolons absent. Leaves monomorphic, evergreen. Petiole ± as long as blade, base not swollen; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade broadly deltate or pentagonal, 2--3-pinnate-pinnatifid, gradually to abruptly reduced distally to pinnate or pinnatifid apex, papery to somewhat leathery. Pinnae not articulate to rachis, segment margins and especially apex spinulose; proximal pinnae largest, petiolulate, inequilateral with basal basiscopic pinnule much larger and more elongate than more distal pinnules; costae adaxially grooved, grooves continuous from rachis to costae to costules; indument of hairlike scales abaxially, absent adaxially. Veins free, forked. Sori in 1 row between midrib and margin, round; indusia round-reniform, attached at narrow sinus, persistent. Spores brownish, rugate or tuberculate, sometimes spiny. x = 41.
Terrestrial ferns. Rhizome creeping or suberect, scaly; scales non-peltate, non-clathrate, mostly entire, brown to red-brown. Fronds tufted or spaced. Stipe scaly. Lamina 2–4-pinnate, broadly deltoid to pentagonal, herbaceous to coriaceous; rachis ridges of the adaxial surface never continuous with the pinna margin; central channel scaly or glabrous. Ultimate segments with or without marginal teeth; veins free. Sori rounded, borne on the vein or at a vein tip; indusium rounded to reniform. Spores monolete, bilateral.
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Growth form herb
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Foliage retention evergreen
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Hardiness (USDA) 7-11

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Cultivation

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Images

Arachniodes unspecified picture

Distribution

Arachniodes world distribution map, present in Australia, China, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:17023720-1
WFO ID wfo-4000002821
COL ID ZKN
BDTFX ID 120241
INPN ID 446400
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Arachniodes Phanerophlebiopsis Leptorumohra Lithostegia Byrsopteris

Lower taxons

Arachniodes x ikeminensis Arachniodes x kenzo-satakei Arachniodes x kurosawae Arachniodes x masakii Arachniodes neopodophylla Arachniodes oohorae Arachniodes quadripinnata Arachniodes x sasamotoi Arachniodes superba Arachniodes x takayamensis Arachniodes tomitae Arachniodes tsiangiana Arachniodes x azuminoensis Arachniodes x clivorum Arachniodes x mitsuyoshiana Arachniodes davalliaeformis Arachniodes blinii Arachniodes bella Arachniodes formosissima Arachniodes sarasiniorum Arachniodes tripinnata Arachniodes aristata Arachniodes simplicior Arachniodes hekiana Arachniodes coniifolia Arachniodes miqueliana Arachniodes mutica Arachniodes cavalerii Arachniodes pseudorepens Arachniodes repens Arachniodes standishii Arachniodes cornucervi Arachniodes palmipes Arachniodes sledgei Arachniodes leucostegioides Arachniodes x minamitanii Arachniodes fengii Arachniodes hunanensis Arachniodes longipinna Arachniodes gigantea Arachniodes amoena Arachniodes globisora Arachniodes hainanensis Arachniodes grossa Arachniodes nipponica Arachniodes x pseudohekiana Arachniodes subamabilis Arachniodes henryi Arachniodes simulans Arachniodes spectabilis Arachniodes chinensis Arachniodes jinpingensis Arachniodes pseudoassamica Arachniodes sinorhomboidea Arachniodes ailaoshanensis Arachniodes rigidissima Arachniodes insularis Arachniodes gansuensis Arachniodes ziyunshanensis Arachniodes x mirabilis Arachniodes tonkinensis Arachniodes sinomiqueliana Arachniodes squamulosa Arachniodes wulingshanensis Arachniodes denticulata Arachniodes miyakei Arachniodes carvifolia Arachniodes x respiciens Arachniodes hiugana Arachniodes rhomboidea Arachniodes assamica Arachniodes pseudoaristata Arachniodes amabilis Arachniodes festina Arachniodes nigrospinosa Arachniodes haniffii Arachniodes puncticulata Arachniodes webbiana