Plants 12-35 cm tall. Rhizome erect, short, apex scaly; scales dark brown to black, narrowly triangular to linear-subulate, margin fimbriate to subentire. Fronds caespitose; stipe shiny, purplish black, 6-15(-18) cm, terete, rigid and threadlike, with brown, multicellular uniseriate hairs or subglabrous; lamina triangular, 7-12 × 5-10 cm, base truncate, bipinnate to tripinnate-pinnatifid, apex acuminate to caudate; pinnae 10-15 pairs, subopposite to alternate, lower pinnae stalked, upper almost sessile, stalk dark brown abaxially, basal 1 or 2 pinnae longest and often falcate, narrowly triangular to oblong, 2.5-7 × 1.5-2 cm, base asymmetrical, on acroscopic side auriculate-truncate, basiscopic side cuneate, up to bipinnate-pinnatifid, apex obtuse; pinnules 6-8 pairs, anadromous, usually sessile but shortly stalked in more divided fronds, basal acroscopic pinnule largest, ovate, 7-9 × 4-7 mm, base asymmetrical, acroscopic side truncate, basiscopic cuneate, apex obtuse to truncate; ultimate segments ovate-oblong to linear, apex with 2-4 short and broadly triangular, obtuse to submucronate or sharp teeth. Costa sulcate adaxially, green, veins often raised adaxially when dry, anadromously branching, (1 or)2-6 veins per ultimate segment, terminal hydathode obvious, and not reaching margin. Frond firmly herbaceous, green, lamina with multicellular uniseriate hairs or subglabrous; rachis shiny purplish black, becoming green in upper half toward apex, with brown, multicellular uniseriate hairs or subglabrous, sulcate adaxially. Sori (1 or)2-4 per pinnule or segment, medial to subterminal on acroscopic veinlets, confluent at maturity, oval to linear, 1-2 mm; indusia grayish green, oval to linear, membranous, repand to entire, opening toward costa or costule, persistent. Spores brown to dark brown, lophate (cristate-alate) perispore. Plants tetraploid: 2n = 144.
Home