Asplenium L.

Spleenwort (en), Asplénie (fr)

Genus

Pteridophytes > Polypodiales > Aspleniaceae

Characteristics

Plants epilithic, epiphytic, or terrestrial. Rhizome rarely dorsiventral, creeping, decumbent, or erect, with clathrate scales. Fronds herbaceous to leathery, sometimes subfleshy, remote or (more often) clustered; stipe dull, green to castaneous or black and then often shiny, base terete or semiterete, often becoming sulcate adaxially, rounded to carinate abaxially; lamina simple to 4-pinnate; rachis sulcate adaxially often with raised central supravascular ridge; margin of pinnae or pinnules often decurrent and forming adaxial wings along rachis or costa; acroscopic side of pinnae, pinnules, and ultimate segments often more developed and these then asymmetrical, pseudodimidiate to dimidiate; margin entire to coarsely serrate, each tooth usually with one veinlet, obtuse, mucronate to acute. Veins free, rarely anastomosing, anadromously branching, rarely with a single vein per pinnule or segment, not reaching margin. Sori linear to subelliptic, rarely double, indusiate; indusia thinly membranous to papery, free margin entire to erose, sinuate or fimbriate, rarely with uniseriate glandular hairs; stalks of sporangia long uniseriate, annuli with 20-28 thickened cells. Spores bilateral, elliptic to reniform, monolete, exospore smooth, perispores elaborate and very variable; in sexual plants 64 spores per sporangium. Plants sexual or agamosporous, x = 35, 36.
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Terrestrial, lithophytic or epiphytic ferns. Rhizome short-to long-creeping, with clathrate scales. Fronds monomorphic. Stipe usually glossy, castaneous to dark purplish or black, rarely grayish green. Lamina simple (not in Australia) or pinnate. Pinnae asymmetric, thin-textured, often with a portion towards the attachment and facing the rhizome (basiscopic) excised; basiscopic margin with main vein following close and parallel near its base, entire; margin facing the apex (acroscopic) crenate or serrate; veins free, rarely anastomosing (not in Australia), conspicuous, usually with 1–many branching acroscopically from main vein of pinna before the first vein branches basiscopically. Sori elongate along veins, away from lamina margins, protected by elongate indusia attached to the veins. Spores bilateral, ellipsoid to subglobose, winged, spiny (not in Australia) or papillose (not in Australia).
Terrestrial, lithophytic or epiphytic ferns. Rhizome erect or short-creeping, rarely long-creeping (not in Australia), with clathrate scales. Fronds monomorphic or rarely dimorphic, sometimes proliferous. Stipe dull or glossy, black, red-brown, brown or green, glabrous to densely hairy or scaly. Lamina simple or 1–5-pinnate, glabrous, hairy or scaly; margin entire to doubly serrate or lobed; pinnae symmetric or asymmetric, thin-textured to thick and fleshy; veins free or rarely anastomosing, inconspicuous or conspicuous. Sori elongate along veins, away from or close to and parallel to lamina margins, protected by elongate indusia attached along veins or indusium absent. Spores echinate (not in Australia), reticulate or costate-alate.
Petiole mostly wiry, green to black, sparsely scaly, otherwise glabrous, with 2 bundles below, these united above into a single bundle mostly X-shaped in section; lvs clustered, the blade simple and entire to thrice pinnate; veins free except in A. rhizophyllum; sori on the lower lf-surface, ± elongate, the indusia attached at one side, all facing distally, thin and hyaline, glabrous, entire; mostly ± evergreen ferns with short-creeping to suberect rhizomes and membranous to coriaceous lvs. 600+, mostly warm reg. (Camptosorus) Our spp. produce ± sterile hybrids and also enter into a large polyploid complex. The following names are believed to apply to sterile hybrids, as indicated:
Indusium well-developed; sporangia and sori as in family. Rhizome short, ascending; stipites clustered; paleae us. conspicuous, hairs us. present. Fronds pinnate to decompound, rarely simple; lamina us. firm to coriac.; veins free or with occ. anastomoses. A subcosmopolitan genus with over 650 spp.
Roots fibrous, not proliferous or proliferous and producing tiny plantlets. Stems erect, rarely long-creeping; scales basally attached, clathrate. Petioles not articulate. Blades 1--4-pinnate, of diverse size and shape. Indusia present. x = 36.
Sori all facing the same way except in a few species which have simple fronds and then the raised line between adjacent sori lacking
Life form -
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality -
Pollination -
Spread -
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.5 - 1.0
Root system fibrous-root rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Mostly in wet forest habitats, but occasionally in rocky sites in alpine and drier regions, including deserts. 
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In very humid or wet sites beside or in creeks and waterfalls among rainforest.
Light 4-6
Soil humidity -
Soil texture -
Soil acidity 2-8
Soil nutriment -
Hardiness (USDA) 8-11

Usage

Some of the Asplenium species are cultivated as ornamentals, particularly A. australasicum (J.Sm.) Hook. and A. ×lucrosum Perrie & Brownsey in Australia. There are species and cultivars suitable for rockeries, vertical gardens, indoors, pots, and those with drooping fronds that are ideal for hanging baskets (see e.g. Elliot & Jones 1982, Jones 1987, Spencer 1995). In inland, highland regions of New Guinea where rainfall is high and salt is scarce, the fronds of A. acrobryum Christ are burnt and the ash is collected and consumed for its salt content (Croft & Leach 1985).
Uses ornamental
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by divisions.
Mode divisions
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Asplenium unspecified picture

Distribution

Asplenium world distribution map, present in Australia, China, New Zealand, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:328226-2
WFO ID wfo-4000003342
COL ID 346X
BDTFX ID 85945
INPN ID 189549
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Arcasplenium Ceterachopsis Pleurosorus Amesium Phyllitis Ceterac Glossopteris Onopteris Phyllitis Rutamuraria Ceterach Diellia Asplenium Camptosorus Trichomanes Acropteris Antigramma Asplenidictyum Biropteris Boniniella Caenopteris Chamaefilix Darea Diplora Eremopodium Holodictyum Micropodium Myriotheca Notolepeum Phyllitopsis Schaffneria Sinephropteris Stormesia Tarachia Thamnopteris Triphlebia Loxoscaphe Neottopteris Scolopendrium

Lower taxons

Asplenium aegaeum Asplenium amaurolobum Asplenium x aran-tohanum Asplenium atuntzeense Asplenium x badense Asplenium x bechereri Asplenium beckeri Asplenium x bouharmontii Asplenium bourgaei Asplenium x brissaginense Asplenium cariocanum Asplenium carnarvonense Asplenium celtibericum Asplenium x centovallense Asplenium x chasmophilum Asplenium comosum Asplenium compressum Asplenium corbariense Asplenium corderoanum Asplenium x cyrnosardoum Asplenium demerkense Asplenium dielerectum Asplenium dimorphum Asplenium domingense Asplenium durum Asplenium x dutartrei Asplenium trindadense Asplenium filiceps Asplenium filidens Asplenium foersteri Asplenium funckii Asplenium gastonis Asplenium x genicoalitum Asplenium glaucostipes Asplenium griseum Asplenium gueinzianum Asplenium hispanicum Asplenium horizontale Asplenium x hungaricum Asplenium x iidanum Asplenium x jacksonii Asplenium jensenii Asplenium x joncheerei Asplenium x kummerlei Asplenium lademannianum Asplenium lambinonii Asplenium laxifolium Asplenium x lellingerianum Asplenium x lessinense Asplenium leucostegioides Asplenium listeri Asplenium x lobmingense Asplenium lolegnamense Asplenium x lucrosum Asplenium macedonicum Asplenium neobrackenridgei Asplenium neohainanense Asplenium newmani Asplenium nitidum Asplenium novo-guineense Asplenium nutans Asplenium x orellii Asplenium paedigens Asplenium platybasis Asplenium plenum Asplenium presolanense Asplenium x protomajoricum Asplenium pseudovulcanicum Asplenium x recoderi Asplenium x reichlingii Asplenium x reichsteinii Asplenium richardii Asplenium x ruscinonense Asplenium x santamariae Asplenium scandens Asplenium scolopendropsis Asplenium sertularioides Asplenium shuttleworthianum Asplenium simile Asplenium x sleepiae Asplenium x sphenocookii Asplenium spirale Asplenium stipicellatum Asplenium stipitiforme Asplenium x stiriacum Asplenium tafanum Asplenium tenuiculum Asplenium terorense Asplenium tosaense Asplenium translucens Asplenium trifoliatum Asplenium trudelii Asplenium x tubalense Asplenium vittaeforme Asplenium wachaviense Asplenium wildii Asplenium haleakalense Asplenium lacinulatum Asplenium muellerianum Asplenium salignum Asplenium acrobryum Asplenium amboinense Asplenium lancifolium Asplenium multiforme Asplenium glanduliserratum Asplenium apoense Asplenium x artanense Asplenium ascensionis Asplenium austrobrasiliense Asplenium x barrancense Asplenium x bavaricum Asplenium belloides Asplenium bicentenniale Asplenium bipinnatifidum Asplenium bradeanum Asplenium burundense Asplenium centrafricanum Asplenium cesatianum Asplenium concolor Asplenium confusum Asplenium x contrei Asplenium creticum Asplenium decompositum Asplenium dognyense Asplenium douglasii Asplenium ekmanii Asplenium elaphoglossoides Asplenium emarginato-dentatum Asplenium campos-portoi Asplenium x joellaui Asplenium cornutissimum Asplenium erythraeum Asplenium exhaustum Asplenium fenzlii Asplenium francii Asplenium gastoni-gautieri Asplenium gibberosum Asplenium gjellerupii Asplenium hapalophyllum Asplenium harleyanum Asplenium harmanii Asplenium hookerianum Asplenium javorkae Asplenium x khaniense Asplenium x krameri Asplenium labillardierei Asplenium ledermannii Asplenium x liberiense Asplenium longissimum Asplenium morobense Asplenium murariaeforme Asplenium mysorense Asplenium x nieschalkii Asplenium octoploideum Asplenium oroupouchense Asplenium pachychlamys Asplenium pacificum Asplenium papuanum Asplenium paradoxum Asplenium parksii Asplenium persicifolium Asplenium pocsii Asplenium ponapeanum Asplenium poscharskyanum Asplenium quezelii Asplenium reuteri Asplenium x sarniense Asplenium schelpei Asplenium seelosii Asplenium setoi Asplenium siamense Asplenium squamuliferum Asplenium staudtii Asplenium subemarginatum Asplenium x suevicum Asplenium x trichomaniforme Asplenium geraense Asplenium tutwilerae Asplenium udzungwense Asplenium x valgannense Asplenium veneticolor Asplenium venustum Asplenium vieillardii Asplenium warburgianum Asplenium werneri Asplenium x wherryi Asplenium balansae Asplenium alvarezense Asplenium aneitense Asplenium anguineum Asplenium anguifrons Asplenium acostae Asplenium rutshuruense Asplenium x kokeense Asplenium x flagrum Asplenium nitens Asplenium kinabaluense Asplenium longum Asplenium scolopendrioides Asplenium loxocarpum Asplenium nephrolepioides Asplenium ruwenzoriense Asplenium stantoni Asplenium stereophyllum Asplenium haughtonii Asplenium glanduliserrulatum Asplenium haurakiense Asplenium heteromorphum Asplenium howeanum Asplenium kelelense Asplenium klossii Asplenium macarenianum Asplenium cymbifolium Asplenium mantalingahanum Asplenium marattioides Asplenium pauperequitum Asplenium perakense Asplenium perlongum Asplenium perplexum Asplenium protractum Asplenium pseudobulbiferum Asplenium pseudocaudatum Asplenium pseudotenerum Asplenium pteridoides Asplenium psychropolitanum Asplenium quaylei Asplenium rapense Asplenium reekmansii Asplenium regis Asplenium sancti-christofori Asplenium sebungweense Asplenium schweinfurthii Asplenium scleroprium Asplenium stenolobum Asplenium schultzei Asplenium setisectum Asplenium sumatranum Asplenium unisorum Asplenium albersii Asplenium bugoiense Asplenium colubrinum Asplenium currorii Asplenium hagenii Asplenium parablastophorum Asplenium x wojaense Asplenium balense Asplenium khullarii Asplenium x capillicaule Asplenium x garhwalense Asplenium rebeccae Asplenium sikkimbirii Asplenium x tandinii Asplenium angolense Asplenium chathamense Asplenium sagittatum Asplenium lacinioides Asplenium schnellii Asplenium difforme Asplenium oblongifolium Asplenium x coto-brusense Asplenium balearicum Asplenium x dolosum Asplenium pagesii Asplenium gregoriae Asplenium poolii Asplenium marojejyense Asplenium paucijugum Asplenium normale Asplenium eylesii Asplenium lividum Asplenium bipartitum Asplenium auritum Asplenium stuhlmannii Asplenium blastophorum Asplenium cuneatum Asplenium viviparioides Asplenium theciferum Asplenium abscissum Asplenium acuminatum Asplenium auriculatum Asplenium bradleyi Asplenium caudatum Asplenium x curtissii Asplenium ebenoides Asplenium exiguum Asplenium feei Asplenium formosum Asplenium gravesii Asplenium x herb-wagneri Asplenium heterochroum Asplenium x heteroresiliens Asplenium horridum Asplenium inexpectatum Asplenium insiticium Asplenium juglandifolium Asplenium kaulfussii Asplenium kentuckiense Asplenium lobulatum Asplenium macraei Asplenium montanum Asplenium myriophyllum Asplenium obtusifolium Asplenium pinnatifidum Asplenium x plenum Asplenium polyodon Asplenium pteropus Asplenium pumilum Asplenium radicans Asplenium serra Asplenium serratum Asplenium sphenotomum Asplenium x stotleri Asplenium x trudellii Asplenium verecundum Asplenium vespertinum Asplenium x wherryi Asplenium antiquum Asplenium praemorsum Asplenium falcatum Asplenium buettneri Asplenium lunulatum Asplenium splendens Asplenium scolopendrium Asplenium dalhousieae Asplenium christii Asplenium lobatum Asplenium preussii Asplenium simii Asplenium macrurum Asplenium dareoides Asplenium trilobum Asplenium africanum Asplenium annetii Asplenium barteri Asplenium brausei Asplenium emarginatum Asplenium hallei Asplenium hemitomum Asplenium jaundeense Asplenium laurentii Asplenium macrophlebium Asplenium megalura Asplenium nigritianum Asplenium subintegrum Asplenium cirrhatum Asplenium alatum Asplenium breedlovei Asplenium sessilifolium Asplenium tuerckheimii Asplenium uniseriale Asplenium virillae Asplenium dissectum Asplenium minimum Asplenium x tyrrhenicum Asplenium pearcei Asplenium yoshinagae Asplenium serricula Asplenium tenuifolium Asplenium acutiserratum Asplenium cuneifolium Asplenium daghestanicum Asplenium lepidum Asplenium punjabense Asplenium capense Asplenium phillipsianum Asplenium azoricum Asplenium petrarchae Asplenium helii Asplenium x adulterinum Asplenium fissum Asplenium ceii Asplenium austrochinense Asplenium capillipes Asplenium coenobiale Asplenium filipes Asplenium griffithianum Asplenium micantifrons Asplenium prolongatum Asplenium ritoense Asplenium tenerum Asplenium bullatum Asplenium phyllitidis Asplenium crinicaule Asplenium x capillipoides Asplenium x chingii Asplenium argentinum Asplenium depauperatum Asplenium biafranum Asplenium subflexuosum Asplenium gomezianum Asplenium excelsum Asplenium maxonii Asplenium escaleroense Asplenium raddianum Asplenium abyssinicum Asplenium variabile Asplenium cornutum Asplenium quitense Asplenium adulterinum Asplenium repens Asplenium barclayanum Asplenium wacketii Asplenium ulbrichtii Asplenium pulchellum Asplenium kunzeanum Asplenium mucronatum Asplenium truncorum Asplenium delicatulum Asplenium purpurascens Asplenium sciadophilum Asplenium x incisoserratum Asplenium projectum Asplenium bulbiferum Asplenium dimidiatum Asplenium adiantifrons Asplenium trapeziforme Asplenium antrophyoides Asplenium fugax Asplenium interjectum Asplenium pseudopraemorsum Asplenium pulcherrimum Asplenium sampsoni Asplenium speluncae Asplenium sublaserpitiifolium Asplenium bifrons Asplenium x mustangense Asplenium x sobtianum Asplenium bradei Asplenium cordatum Asplenium holosorum Asplenium grevillei Asplenium eatonii Asplenium oligosorum Asplenium underwoodii Asplenium nesioticum Asplenium dayi Asplenium rhomboidale Asplenium diplosceum Asplenium jenmanii Asplenium clutei Asplenium scortechinii Asplenium callipteris Asplenium paucivenosum Asplenium fibrillosum Asplenium biscaynianum Asplenium palmeri Asplenium herpetopteris Asplenium cancellatum Asplenium subaequilaterale Asplenium kassneri Asplenium lastii Asplenium lokohoense Asplenium sandersonii Asplenium tsaratananense Asplenium mangindranense Asplenium andapense Asplenium anisophyllum Asplenium diplazisorum Asplenium boiteaui Asplenium correardii Asplenium protensum Asplenium mildbraedii Asplenium apertum Asplenium afzelii Asplenium mannii Asplenium leandrianum Asplenium parvisorum Asplenium dregeanum Asplenium thunbergii Asplenium virchowii Asplenium clermontiae Asplenium contiguum Asplenium rutaceum Asplenium schizophyllum Asplenium claussenii Asplenium australasicum Asplenium flaccidum Asplenium annobonense Asplenium macrosorum Asplenium obtusatum Asplenium triphyllum Asplenium x boydstoniae Asplenium x shawneense Asplenium gemmiferum Asplenium castaneum Asplenium flabellulatum Asplenium harpeodes Asplenium harrisii Asplenium miradorense Asplenium polyphyllum Asplenium obesum Asplenium otites Asplenium hemionitis Asplenium hastatum Asplenium squamosum Asplenium hostmannii Asplenium ambohitantelense Asplenium bangii Asplenium ecuadorense Asplenium oellgaardii Asplenium pseudoangustum Asplenium gilbertii Asplenium tadei Asplenium tripteropus Asplenium flexuosum Asplenium zamiifolium Asplenium stuebelianum Asplenium gemmascens Asplenium holophlebium Asplenium pululahuae Asplenium novae-caledoniae Asplenium oligolepidum Asplenium purdieanum Asplenium holstii Asplenium linckii Asplenium davisii Asplenium rosenstockianum Asplenium aureum Asplenium sulcatum Asplenium congestum Asplenium virens Asplenium canelense Asplenium volkensii Asplenium poloense Asplenium oblongatum Asplenium potosinum Asplenium microtum Asplenium nesii Asplenium ghiesbreghtii Asplenium delavayi Asplenium lorentzii Asplenium formosae Asplenium cuneatiforme Asplenium sylvaticum Asplenium vulcanicum Asplenium hallbergii Asplenium stolonipes Asplenium yelagagense Asplenium kiangsuense Asplenium ruizianum Asplenium erosum Asplenium x fawcettii Asplenium scandicinum Asplenium actiniopteroides Asplenium loxoscaphoides Asplenium torrei Asplenium pringlei Asplenium haenkeanum Asplenium vomeriforme Asplenium peruvianum Asplenium discrepans Asplenium tabinense Asplenium herzogii Asplenium brasiliense Asplenium recumbens Asplenium simplicifrons Asplenium oligophyllum Asplenium pseudonitidum Asplenium mourai Asplenium sellowianum Asplenium cruegeri Asplenium xinyiense Asplenium barkamense Asplenium x wangii Asplenium hainanense Asplenium saxicola Asplenium pseudovarians Asplenium pedicularifolium Asplenium adamsii Asplenium achalense Asplenium arcanum Asplenium goetzii Asplenium trapezoideum Asplenium cimmeriorum Asplenium procerum Asplenium eberhardtii Asplenium semipinnatum Asplenium carvalhoanum Asplenium mossambicense Asplenium ramlowii Asplenium chaseanum Asplenium pseudoauriculatum Asplenium kenzoi Asplenium eutecnum Asplenium gentryi Asplenium tryonii Asplenium ultimum Asplenium venturae Asplenium rockii Asplenium humbertii Asplenium uhligii Asplenium paucifolium Asplenium smedsii Asplenium ayopayense Asplenium bolivianum Asplenium micropaleatum Asplenium mosetenense Asplenium tunquiniense Asplenium flabellifolium Asplenium jahandiezii Asplenium drepanophyllum Asplenium trilobatum Asplenium palaciosii Asplenium sessilipinnum Asplenium vareschianum Asplenium badinii Asplenium flaccidum Asplenium milnei Asplenium surrogatum Asplenium hobdyi Asplenium regulare Asplenium nigrescens Asplenium anogrammoides Asplenium simonsianum Asplenium oldhamii Asplenium viviparum Asplenium fujianense Asplenium lilloanum Asplenium x ticinense Asplenium x papyraceum Asplenium malcolm-smithii Asplenium monanthes Asplenium x sollerense Asplenium souchei Asplenium indusiatum Asplenium x kidoi Asplenium lepturus Asplenium athyrioides Asplenium caripense Asplenium ocoense Asplenium mauritiensis Asplenium xianqianense Asplenium martianum Asplenium x uiryeongse Asplenium blepharodes Asplenium dianae Asplenium modestum Asplenium pifongiae Asplenium hypomelas Asplenium cristatum Asplenium elliottii Asplenium erectum Asplenium inaequilaterale Asplenium salicifolium Asplenium affine Asplenium rutifolium Asplenium x alternifolium Asplenium salwinense Asplenium komarovii Asplenium nidoides Asplenium pleurosorum Asplenium subspathulinum Asplenium attenuatum Asplenium rectangulare Asplenium x ligusticum Asplenium x bosco-gurinense Asplenium boltonii Asplenium costei Asplenium elmeri Asplenium isabelense Asplenium x kitazawae Asplenium x mendelianum Asplenium x rosselloi Asplenium rukaraense Asplenium simplicifolium Asplenium tamandarei Asplenium x tavelio Asplenium x waikamoi Asplenium warneckii Asplenium biscayneanum Asplenium solmsii Asplenium chimantae Asplenium cowanii Asplenium altajense Asplenium sarelii Asplenium sajanense Asplenium atroviride Asplenium sanchezii Asplenium pellucidum Asplenium daucifolium Asplenium wrightii Asplenium macrophyllum Asplenium stoloniferum Asplenium stellatum Asplenium incisum Asplenium finlaysonianum Asplenium x virginicum Asplenium trigonopterum Asplenium laserpitiifolium Asplenium oligophlebium Asplenium ensiforme Asplenium castaneoviride Asplenium extensum Asplenium fragrans Asplenium longicauda Asplenium petiolulatum Asplenium laciniatum Asplenium prionites Asplenium x shawneense Asplenium tenerrimum Asplenium blepharophorum Asplenium barbaense Asplenium cuspidatum Asplenium monodon Asplenium hybridum Asplenium hallii Asplenium tricholepis Asplenium lamprocaulon Asplenium gilliesii Asplenium x morganii Asplenium munchii Asplenium shikokianum Asplenium x mitsutae Asplenium cladolepton Asplenium pseudoerectum Asplenium divaricatum Asplenium aequibasis Asplenium x akaishiense Asplenium amoenum Asplenium araucarieti Asplenium athertonense Asplenium atrovirens Asplenium batuense Asplenium bicarinatum Asplenium x claphamii Asplenium clemensiae Asplenium cromwellianum Asplenium x diasii Asplenium dielpallidum Asplenium x tagananaense Asplenium erythrocaulon Asplenium eurysorum Asplenium gardneri Asplenium impressivenium Asplenium x inexpectatum Asplenium lamprophyllum Asplenium lobangense Asplenium longipes Asplenium lyallii Asplenium monotis Asplenium paucidens Asplenium peteri Asplenium picardae Asplenium dichotomum Asplenium dicranurum Asplenium dielfalcatum Asplenium dielmannii Asplenium diplotion Asplenium disjunctum Asplenium x mantoniae Asplenium perardii Asplenium seileri Asplenium ruprechtii Asplenium wilfordii Asplenium microxiphion Asplenium x murbeckio Asplenium musifolium Asplenium parvum Asplenium schizocarpum Asplenium schizotrichum Asplenium shawii Asplenium shimurae Asplenium steerei Asplenium subaquatile Asplenium tenuicaudatum Asplenium woynarianum Asplenium zenkerianum Asplenium powellii Asplenium haenkei Asplenium kjellbergii Asplenium borneense Asplenium brachycarpum Asplenium capitisyork Asplenium carinatum Asplenium carruthersii Asplenium praegracile Asplenium dentatum Asplenium insolitum Asplenium olivaceum Asplenium soleirolioides Asplenium sphaerosporum Asplenium decrescens Asplenium adnatum Asplenium asterolepis Asplenium resiliens Asplenium obovatum Asplenium rhizophyllum Asplenium nidus Asplenium platyneuron Asplenium aethiopicum Asplenium marinum Asplenium ruta-muraria Asplenium fontanum Asplenium adiantum-nigrum Asplenium viride Asplenium septentrionale Asplenium trichomanes Asplenium onopteris Asplenium ceterach