Asplenium ruta-muraria L.

Wallrue (en), Rue des murailles (fr), Doradille rue-des-murailles (fr), Rue-des-murailles (fr)

Species

Pteridophytes > Polypodiales > Aspleniaceae > Asplenium

Characteristics

Plants 2-10(-15) cm tall. Rhizome erect to shortly creeping, scaly; scales blackish brown, narrowly triangular, occasionally with marginal glands, glands also on apex, subentire. Fronds caespitose; stipe (0.5-)2-5(-6) cm, green or grayish green to stramineous when dry, base castaneous to dark purplish or blackish brown, with numerous short (4-cellular) glandular hairs and hairlike scales, subglabrous toward rachis; lamina deltoid-triangular to ovate, (1-)2-5(-8) × 1-3(-5) cm, apex obtuse to subacute, 2-pinnate or pinnate-pinnatifid; pinnae 1-4 pairs, subopposite or alternate, basal pinnae largest, (4-)8-12 × (4-)8-10 mm, stalked, pinnae triangular, apex obtuse, imparipinnate or ternate with up to 3 lateral segments, segments often similarly divided; ultimate segments broadly flabellate to rhombic-ovate or trapeziform, (3-)4-6 × 2-5 mm, base broadly cuneate and decurrent on costa, margin irregularly dentate, apex obtuse. Veins obscure, flabellately anadromous, and almost parallel. Fronds subleathery, grayish green when dry; rachis and costa green, with (partly) deciduous glandular hairs, subglabrous when old, adaxially sulcate, when dry often flat. Sori 5-12 per pinna, median on subtending vein, confluent at maturity, subelliptic to linear, 1-5 mm; indusia gray-glaucous to brown, linear, thinly membranous, margin with long hairs, opening toward major veins or costa, concealed by sporangia at maturity. Perispore lophate (costate-cristate), average exospore length 41-49 µm. Plants sexual, autotetraploid: 2n = 144.
More
Roots not proliferous. Stems short-creeping to erect, often branched; scales very dark brown throughout, narrowly deltate, 1--3 × 0.1--0.25 mm, margins with widely spaced teeth. Leaves monomorphic. Petiole reddish brown proximally, green distally, dull, 1--9 cm, (1/2--)1--2 times length of blade; indument of dark brown, narrowly deltate scales proximally grading into multicellular hairs. Blade deltate-ovate to obovate or oblanceolate, 1--2(--3)-pinnate to 2-pinnate-pinnatifid, 2--6 × 1--4 cm, somewhat thick, glabrous; base obtuse; apex acute to rounded, not rooting. Rachis green, dull, glabrous except for very sparse, minute hairs. Pinnae in 2--4 pairs, deltate-ovate to obdeltate; proximal (largest) pinnae 7--30 × 5--20 mm; base truncate to acute; margins finely (sometimes coarsely) incised; apex rounded to acute. Veins free, evident. Sori as many as 30 or more per pinna, usually 1--5 per segment, on both basiscopic and acroscopic sides. Spores 64 per sporangium. 2 n = 144.
Rhizome 1.5–2 mm thick, the lanceolate scales 1.5–4 mm; lvs numerous, spreading, 4–17 cm, the petiole to 7 cm, green blade ovate or rhombic, firm, bipinnate or bipinnate-pinnatifid below, merely pinnate above, the rachis green; pinnae alternate, ca 6 pairs, long-petiolulate (petiolule 2–5 mm), the basal ones with 3–6 flabellate or rhombic, finely toothed and sometimes deeply lobed pinnules to 12 mm; 2n=144. Rock-crevices and ledges, mostly on limestone; interruptedly circumboreal, in Amer. from Mass. and Vt. to Mich., s. to Ala., Ill., Mo., and Ark. (A. cryptolepis, the Amer. pls.)
A fern.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention evergreen
Sexuality monoecy
Pollination hydrogamy
Spread anemochory
Mature width (meter) 0.15
Mature height (meter) 0.05 - 0.1
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway -

Environment

Old walls and basic rocks in hilly areas.
More
It is a temperate plant.
Light 3-7
Soil humidity 3-7
Soil texture 4-9
Soil acidity 6-9
Soil nutriment -
Hardiness (USDA) 5-8

Usage

Uses medicinal
Edible fronds
Therapeutic use Expectorant (unspecified), Medicine (unspecified), Pectoral (unspecified), Swelling (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -23
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Asplenium ruta-muraria habit picture by beurny (cc-by-sa)
Asplenium ruta-muraria habit picture by beurny (cc-by-sa)
Asplenium ruta-muraria habit picture by 🐞Hélène🐦 (cc-by-sa)

Leaf

Asplenium ruta-muraria leaf picture by Olivier Eudes (cc-by-sa)
Asplenium ruta-muraria leaf picture by leptospermum scoparium (cc-by-sa)
Asplenium ruta-muraria leaf picture by Martin Allan (cc-by-sa)

Flower

Asplenium ruta-muraria flower picture by Michel AMBROISE (cc-by-sa)

Fruit

Asplenium ruta-muraria fruit picture by 🐞Hélène🐦 (cc-by-sa)
Asplenium ruta-muraria fruit picture by hieracium eriophorum (cc-by-sa)
Asplenium ruta-muraria fruit picture by Yoan MARTIN (cc-by-sa)

Distribution

Asplenium ruta-muraria world distribution map, present in China, France, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:311563-2
WFO ID wfo-0001109353
COL ID HLD8
BDTFX ID 7681
INPN ID 84521
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Amesium germanicum Amesium ruta var. muraria Asplenium zoliense Tarachia multicaulis Tarachia ruta-muraria Belvisia ruta-muraria Scolopendrium ruta var. muraria Tarachia germanica Trichomanes breynii Trichomanes ruta var. muraria Asplenium matthioli Asplenium murorum Chamaefilix ruta-muraria Asplenium subtenuifolium Phyllitis ruta var. muraria Acrostichum ruta var. muraria Asplenium multicaule Asplenium ruta-muraria var. subtenuifolium Asplenium ruta-muraria var. subtenuifolium Asplenium ruta-muraria

Lower taxons

Asplenium ruta-muraria var. cryptolepis Asplenium ruta-muraria var. eberlei Asplenium ruta-muraria subsp. dolomiticum Asplenium ruta-muraria var. lanceolum Asplenium ruta-muraria var. schriesheimense