Plants terrestrial or epilithic. Rhizome short-creeping, sparsely branched, to 35 mm long, to 4 mm in diameter, set with roots, closely spaced persistent stipe bases and scales, scales chartaceous, atrocastaneous, clathrate, sessile, subulate, cordate-imbricate, entire, apex terminates in an oblong thin-walled cell, to 7 mm long, to 1 mm wide. Fronds crowded, caespitose, erect to suberect, to 340 mm long; stipe firm, atrocastaneous, adaxially sulcate, to 146 mm long, to 1.5 mm in diameter, initially moderately scaled, scales chartaceous, atrocastaneous to ferrugineous, clathrate, sessile, subulate to filiform, cordate to cordate-imbricate, with short and long filiform outgrowths at the base, apex entire, terminating in an oblong thin-walled cell, to 9 mm long, to 0.6 mm wide; lamina anadromous, to 2-pinnate, oblong-acute, to 195 mm long, to 50 mm wide, with up to 15 pinna pairs; rachis firm, proximally castaneous, greenish towards the apex, adaxially sulcate, moderately scaled, scales similar to those on the stipe; pinnae petiolate, petiole to 2.5 mm long, opposite to alternate, basally widely spaced, more closely spaced towards the apex and slightly overlapping, firmly herbaceous, rhomboid to trullate, to 30 mm long, to 21 mm wide, usually with a single pinnule pair; pinna-rachis sulcate adaxially, sulcus confluent with that of the rachis, moderately set with scales similar to, but smaller than, those on the rachis; pinnules narrowly flabellate, rhomboid, or oblong, the larger divided into oblong lobes, irregularly dentate, to 13 mm long, to 6 mm wide, adaxially sparsely set with filiform clathrate scales, sessile, filiform, with short and long filiform outgrowths at the base, apex entire, to 2.5 mm long. Venation obscure, flabellate, terminating in the teeth near the margin. Sori linear, to 6 mm long; indusium firmly herbaceous, stramineous, linear, entire, to 6 mm long, to 0.3 mm wide; sporangium long-stalked, uniseriate, 3-seriate below capsule, capsule broadly elliptic in lateral view, with (17-)19(-20) indurated annulus cells, epistomium 2(-3)-celled, hypostomium 4-celled. Spores 64 per sporangium, brown, elliptic, monolete, perispore with low, closely spaced, reticulate ridges, (38-)41.5(-44) x (24-)25.1(-28) µm.
Home