Plants 20-40 cm tall. Rhizome erect, short, scaly; scales blackish brown, narrowly triangular to deltoid-tridentate with long apical tail. Fronds caespitose; stipe grayish brown to purplish black, 5-12 cm, semiterete, adaxially grooved, with blackish brown deltoid-tridentate scales with long apical tail; lamina lanceolate to linear-lanceolate, 10-30 × 3.5-8 cm, gradually reduced at base, apex acute, 1-pinnate; pinnae 18-30(-40) pairs, alternate or lower ones (sub)opposite, almost sessile to shortly stalked; basal pinnae slightly reduced, deltoid-triangular to ovate, apex obtuse; middle pinnae narrowly subtriangular to rhomboid-elliptic, 2-4(-5.5) × 0.8-1.5 cm, base asymmetrical, acroscopic side truncate and auriculate, basiscopic side narrowly cuneate, margin irregularly biserrate-crenate, apex acute. Veins obvious, mostly 2-forked, 3-forked, or simple, not reaching margin. Fronds papery, dark brown when dry, both surfaces with dark brown stellate or hairlike small scales, subglabrous when old; rachis grayish brown, semiterete, adaxially grooved, with rigid hairlike scales, rarely with subapical gemma. Sori linear to crescent-shaped, 4-8 mm, on acroscopic veinlets; indusia yellowish brown to grayish brown, linear, membranous, entire, opening toward costa or to first acroscopic vein, often rolling back at maturity, persistent. Perispore costate, with average exospore length 40-44 (dodecaploids). Plants sexual polyploid: 2n = 144, 288, or 432.
Home