Plants terrestrial or epilithic. Rhizome erect to suberect, to 35 mm long, to 3 mm in diameter, set with roots, crowded persistent stipe bases and scales, scales chartaceous to crustaceous, castaneous, clathrate, sessile, lanceolate, cordate to cordate-imbricate, with scattered filiform outgrowths along the margin of which the apex terminates in an elliptic thin-walled cell, apex terminates in an oblong thin-walled cell, to 8 mm long, to 1 mm wide. Fronds crowded, caespitose, erect to arching, to 510 mm long; stipe firm, castaneous, matt, terete, to 120 mm long, to 1.8 mm in diameter, narrowly green-winged in the upper half, initially moderately set with ferrugineous glandular cells and filiform clathrate scales of which the apex terminates in an enlarged gland-like cell, to 3 mm long, glabrous later; lamina 1-pinnate, narrowly elliptic to oblong-cuneate, to 390 mm long, to 60 mm wide, with up to 36 stalked pinna pairs; rachis firm, castaneous, terete, narrowly green-winged, initially moderately set with scales similar to those on the stipe, glabrous later; pinnae sessile, herbaceous, opposite to alternate, basally more widely spaced than apically, pinnae in the lower half to one third of the lamina gradually reduced, inequilaterally lanceolate to ovate, often slightly auricled acroscopically, acroscopically truncate to broadly cuneate, basiscopically narrowly to broadly cuneate, dentate, to 30 mm long, to 14 mm wide, glabrous adaxially and abaxially; costa adaxially raised, convex. Venation anadromous, evident, pinnately branched, branches forked, ending in the teeth near the margin. Sori linear, inframedial, usually below or at a vein fork, to 3.5 mm long; indusium chartaceous, stramineous, linear, entire, to 3.5 mm long, to 0.3 mm wide; sporangium long-stalked, uniseriate, 3-seriate below the capsule, capsule globose in lateral view, annulus with (17-)19(-21) indurated annulus cells, epistomium 2-celled, hypostomium (3-)4-celled. Spores 64 per sporangium, brown, elliptic, monolete, perispore forms broad reticulate wings, exospore (26-)28.8(-38) x (16-)18.5(-24) µm.
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Rhizome erect to suberect, up to 35 mm long, up to 3 mm in diam., with crowded stipe bases and castaneous, lanceolate scales, up to 8 x 1 mm, with scattered filiform outgrowths along margins. Fronds caespitose, erect to arching, up to 510 mm long. Stipe firm, castaneous, up to 120 mm long, up to 1.8 mm in diam., narrowly green-winged in upper half, with glandular cells and ferruginous, filiform scales, up to 3 mm long, glabrous later. Lamina 1-pinnate, narrowly elliptic to oblong-cuneate, up to 390 x 60 mm, with up to 36 petiolate pinna pairs. Rachis firm, castaneous, terete, narrowly green-winged. Pinnae herbaceous, inequilaterally lanceolate to ovate, often slightly auricled acroscopically, basiscopically narrowly to broadly cuneate, dentate, up to 30 x 14 mm, glabrous adaxially and abaxially. Sori linear, up to 3.5 mm long. Indusium linear, entire, up to 3.5 x 0.3 mm.
Rhizome erect, c. 3 mm in diameter, set with dark brown, lanceolate, acuminate, subentire rhizome-scales c. 5 mm long. Fronds tufted, firmly membranous, not gemmiferous; stipe castaneous, usually less than 1/4 lamina length, with narrow green wings in upper half and sparsely scaly at first; lamina narrowly elliptic, up to 400 x 55 mm, pinnate to 2-pinnatifid, winging rhachis, lower pinnae decrescent; pinnae in up to 40 pairs, somewhat falcate, acroscopic basal lobe separated to a greater or lesser extent, base unequally cuneate, margin deeply serrate, serrations single or paired, glabrous. Sori up to 15 per pinna, extending from near costa 1/3-2/3 to margin; indusium membranous, entire to erose, c. 0.3 mm broad.
Perennial with short, erect or suberect rhizome. Fronds to 51 cm, erect to arching, stipe subglabrous, narrowly green-winged in upper half, lamina 1-pinnate to 2-pinnatifid, narrowly elliptic, pinnae to 3 cm long, falcate, obtusely dentate. Sori to 15 per pinna, indusium entire to erose.
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