Roots not proliferous. Stems short-creeping to erect, frequently branched; scales dark reddish brown, narrowly deltate, 3--5 × 0.3--0.5 mm, margins entire. Leaves monomorphic. Petiole dark reddish brown at base, fading to green in distal 1/3--1/2, lustrous, 1--10 cm, 1/5--1 times length of blade; indument of dark reddish brown, narrowly deltate scales at very base, grading distally into hairs. Blade narrowly deltate, often irregular in outline, pinnatifid or often with single pair of pinnae proximally, 2--17(--20) × 1--4(--13) cm, thick, pubescent abaxially only; base truncate, cordate, or auriculate; apex acute to long-attenuate, proliferous bud very rare, not known to root in nature. Rachis green, sometimes drying to tan, dull; hairs on abaxial surface only, scattered, minute. Pinnae 0--1 pair, ovate to deltate, sometimes narrowly so, 5--20(--90) × 0.4--1(--1.2) mm; base truncate to acute; margins crenate to serrate; apex rounded to attenuate. Veins free (rarely anastomosing), obscure. Sori 1--6(--40+) per segment, usually confluent with age. Spores 64 per sporangium. 2 n = 144.