Annuals; not scapose; glabrous or pubescent, trichomes simple mixed with short-stalked, forked, 3-rayed, or cruciform ones. Stems (few from base), ascending, branched. Leaves basal and cauline; petiolate or sessile; basal not rosulate, shortly petiolate, blade margins entire or dentate; cauline shortly petiolate or sessile, blade (base not auriculate), margins entire or dentate. Racemes (corymbose, several-flowered, secund, lax), considerably elongated in fruit. Fruiting pedicels recurved or reflexed, slender or stout, (glabrous or pubescent). Flowers (cleistogamous and/or chasmogamous); sepals suberect, oblong, lateral pair not saccate basally, (glabrous or pubescent); petals white, (usually rudimentary, or well-developed and exceeding sepals, rarely absent), spatulate or oblong, claw indistinct; stamens subequal; filaments slightly dilated or not basally; anthers ovate to globose; nectar glands lateral, each side of lateral stamens. Fruits (pendulous), silicles, indehiscent or very tardily dehiscent, sessile, usually orbicular or obovate to elliptic, rarely oblong, twisted or flattened, latiseptate; valves each not veined or veins prominent, pubescent or glabrous; replum rounded; septum absent or complete; ovules 2-12 per ovary; style obsolete or distinct; stigma capitate. Seeds uniseriate, flattened, not winged, oblong; seed coat not mucilaginous when wetted; cotyledons accumbent.