Small to large trees, ultimate shoots with complicated system of divaricate branching. Stipules large, prominently keeled, lateral, persistent or subpersistent. Leaves almost glabrous on both surfaces, often with minute papillae on venation giving beaded appearance, without stomatal crypts, with a pair of glands on midrib at or near base of lower surface. Petioles eglandular. Inflorescence a raceme, or sparsely branched, contracted panicle. Bracts and bracteoles persistent, eglandular, not enclosing groups of flower buds. Flowers hermaphrodite. Receptacle obconical to cylindrical, as long as or exceeding calyx lobes, hollow, hairy inside throughout, throat blocked by retrorse hairs. Calyx lobes 5, broadly ovate to lanceolate, tomentellous on both surfaces. Petals 5, glabrous, exceeding calyx lobes. Stamens 10-20, posterior, inserted unilaterally on margin of disk; filaments free, exserted; staminodes forming a barely visible denticulate margin to throat. Ovary inserted at mouth of receptacle tube, pilose on exterior; carpel bilocular with 1 ovule in each loculus. Fruit large; epicarp glabrous, densely verrucose-crustaceous; mesocarp transversely fibrous; endocarp hard, thick, shortly and sparsely hairy inside, breaking up irregularly at germination. Cotyledons large and strongly ruminate. Germination cryptocotylar, eophylls alternate.
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