Batis P.Browne

Turtleweed (en)

Genus

Angiosperms > Brassicales > Bataceae

Characteristics

Glabrous, monoecious or dioecious shrubs, often nodding or prostrate; the bark flaking off; branching alternate, sometimes rooting and branching at the nodes. Leaves opposite, succulent, linear, apically pointed or mucronate, basally clasping but drying with an umbonate appendage; stipules inconspicuous, caducous. Inflorescences axillary or terminal, in monoecious species the flowers borne separately on short leafy axes (brachyblasts) with male and female flowers sessile on the same shoot, in dioecious species the flowers borne in short, dense axillary catkins or syncarps. Male flowers subtended by a bract or leaf and sur-rounded by a pair of tepallike bracts (spathella), the stamens 4, alternating with 4 staminodes or appendages, the anthers ovoid, exserted, the filaments glabrous, the gynoecium rudimentary or wanting; female flowers or catkins subtended by a pair of scalelike bracts or by leaves, the pistils fused or separate, when fused bearing a scalelike bract near the apex, the pistil 4-locular, each locule with a single erect, basal ovule, the style persistent, short or wanting, the stigmas 2-capi-tate, fimbriate, persistent. Fruits drupes or syncarps, the outer tissues fleshy or leathery, the endocarp woody; seeds 1-4 in each pistil, oblong, nearly straight.
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Dioecious or monoecious small shrubs with thick woody roots. Leaves simple, opposite, sessile, fleshy, with a distinctly saccate, colourless base. Stipules minute. Flowers unisexual, either solitary and terminal or axillary, or in small axillary spikes. ♂ Flowers subtended by bracts, enclosed in a membranous spathella which opens with one or two transverse or radial slits giving rise to 2-4 lobes. Tepals 4, valvate. Stamens 4, alternitepalous; anthers dorsifixed, introrse, dehiscing lengthwise with 2 slits. Sometimes an abortive gynaecium present. ♀ Flowers merely consisting of a naked ovary, in the axil of leaves when solitary, in the axil of cordate bracts when growing in spikes, 2-carpellate, 4-celled by one true and one false septum; ovules 1 in each cell, basal, anatropous, with a long funicle. Stigmas 2, sessile, distinctly papillate. Fruit a septicidal berry dehiscing with 2 valves, either solitary or many united together with the bracts into a connate, spikelike whole. Seeds with a large, straight embryo, exalbuminous.
Plants relatively low, sprawling. Leaf blades obovoid to oblanceoloid. Spikes subsessile, ellipsoid, subglobose, or turbinate [lax, bracteate, flowers solitary]. Flowers anemophilous; filaments slender, or sometimes winged; anthers versatile, dorsifixed; stigmas sessile, papillate. Syncarps each with 1-4 seeds (pyrenes). Seeds narrow, flattened; coats thin. x = 11.
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Environment

The Malaysian species, B. argillicola is found on littoral,rarely flooded, saline clay plains around Merauke in southern New Guinea, behind the mangroves where it occurs sometimes gregariously over large areas though it is sometimes also found solitary on the edge of the mangroves and of bushes scattered over the plains. It seems not to be entirely restricted to clayey soils as it is also found on sandy banks along small creeks in open connection with the sea but out of reach of normal high tide. For a full description of the habitat of this species cf. van Royen Nova Guinea n.s. 1 1956 176-180 f. 1, t. 9. Fl. Aug.-Oct. (end dry season), fr. Dec.-Jan. (wet season).B. maritima, the American species, is also found on saline clay plains, but these appear to be regularly flooded by the tides.
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Hardiness (USDA) 5-12

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Images

Batis unspecified picture

Distribution

Batis world distribution map, present in Australia, Panama, Papua New Guinea, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:30121228-2
WFO ID wfo-4000004220
COL ID 38T3
BDTFX ID -
INPN ID 627874
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Batis

Lower taxons

Batis maritima Batis argillicola