Trees, dioecious. Leaves alternate and distichous; stipules lateral, free; blade entire, pinnately veined. Inflorescences in the leaf axils, pedunculate, with bracts. Staminate inflorescences spicate; tepals 3-5, free; stamens 1 or 2; pistillode absent. Pistillate inflorescences globose-capitate; perianth tubular, subentire to 4-lobed; ovary free, stigmas 2, subulate. Fruiting perianth enlarged, fleshy, greenish; fruit free; seed large, without endosperm, cotyledons thick and equal.