Submerged aquatic herbs, sometimes rooting at the nodes, glabrate or pubes-cent, glandular punctate. Leaves verticillate, pinnately dissected, flat, the seg-ments filiform. Inflorescences solitary flowers in the leaf axils, the pedicels elongating in fruit, ebracteate. Flowers with the calyx divided to near the base into 5 subequal narrow lobes; corolla 2-lipped, the lobes rounded; stamens 4, didynamous, the anthers equal, the thecae equal and proximal; ovary obtuse, 2-locular, the stigma flat and curved. Capsule ovoid, membranaceous, the seeds oblong, longitudinally reticulate, numerous.