Herbs erect, perennial, pilose. Basal leaves long petiolate; leaf blade broadly ovate to lanceolate, base strongly cordate, margin coarsely crenate. Verticillasters many flowered, in terminal spikes; floral leaves sessile; bracteoles at least as long as calyx. Calyx tubular-campanulate, 5-veined, hairy, glabrous at throat inside; teeth 5, equal, straight, rigidly spinescent. Corolla cylindric, at least as long as calyx, 2-lipped, without hairy annulus inside, straight or slightly curved downward, rarely dilated at throat; upper lip concave, margin entire or emarginate; lower lip spreading, 3-lobed; middle lobe longer than lateral lobes, obtuse to emarginate. Stamens 4, parallel, ascending to underside of upper lip of corolla; anterior stamens longer, rarely curved laterally at throat after anthesis; anther cells 2, subparallel. Style apex subequally 2-cleft. Nutlets rounded to subtruncate.
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