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Trees, evergreen, perennial, dioecious or rarely monoecious; stems and foliage without latex. Indumentum of simple, multicellular trichomes; glandular trichomes absent; stinging trichomes absent. Stipules entire, inconspicuous, deciduous. Leaves alternate, petiolate, trifoliate, denticulate to crenate, penniveined; glands absent. Inflorescences axillary, paniculate, solitary, unisexual, with flowers in bracteate clusters. Male flowers pedicellate; calyx lobes 5, valvate, free; petals absent; disc absent; stamens 5, with filaments shortly connate; anthers dorsifixed, bilobate, with thecae oblong and longitudinally dehiscent; pistillodes short stipitate, peltate. Female flowers pedicellate; calyx lobes 5, imbricate, free; petals absent; disc absent; ovary 3-locular; ovules biloculate; styles 3, free, simple. Fruits baccate, subglobose, surface smooth, indehiscent and fleshy. Seeds subglobose; testa crustaceous; albumen fleshy; ecarunculate; cotyledons broad, flat.
Trees, usually dioecious, rarely monoecious, hairy, becoming glabrate, hairs simple; branching not phyllanthoid. Leaves deciduous, alternate, pinnately compound, leaflets 3(or 5), all well developed; stipules fugacious; blade margins crenate-serrate. Inflorescences unisexual, racemose or paniculate thyrses. Pedicels present. Staminate flowers: sepals 5, distinct; petals 0; nectary absent; stamens 5; filaments distinct, adnate to pistillode base; connectives not extending beyond anthers; pistillode short-stipitate, peltate, 5-angled. Pistillate flowers: sepals deciduous, 5, distinct; petals 0; nectary absent; pistil 3(–4)-carpellate; styles 3(–4), connate at base, unbranched. Fruits drupes. Seeds 1–2 per locule, rounded-trigonous; seed coat dry, smooth; caruncle absent. x = 14.
