Trees [shrubs], trunks often swollen proximally, plants monoecious. Leaves: stipules caducous [persistent]; blade often lobed, base usually cuneate to truncate or cordate, margins usually entire or serrate, palmately or pinnately veined. Flowers functionally unisexual; sepals ± valvate, basally connate; nectaries 0 or 10–16; stamens (staminodes in pistillate flowers) [10–]25–30; anthers often crowded; ovary stalked, carpels 5, distinct; styles 5, connate; stigmas 5. Follicles woody, outsides glabrous, insides stellate-hairy. Seeds hairy. x = 20.