Shrubs or trees, usually prickly and tomentose. Leaves digitately compound, palmately lobed or simple, with a sheathing base and a usually bicuspid ligule; petiole terete. Inflorescence usually a terminal panicle of umbellules, often with persistent small bracts; pedicel not articulated below the ovary. Calyx rim 5-den-tate. Petals 5, valvate. Stamens 5, extrorse. Ovary inferior, 2-celled; disk shallow; styles united into a usually long column. Fruit globose, exocarp fleshy, endocarp chartaceous; seed 2, not compressed, endosperm smooth.