Bromus diandrus Roth

Ripgut brome (en), Brome à deux étamines (fr), Anisanthe à deux étamines (fr)

Species

Angiosperms > Poales > Poaceae > Bromus

Characteristics

Annuals, 20-90 cm, loosely tufted or consisting of a solitary culm. Leaf-sheath with soft, short, spreading hairs; upper sheaths ± glabrous near base. Ligule 1.5-4 mm, denticulate to lacerate. Leaf-blade 4-30 cm × 2-7 mm, with short, ± scattered hairs, scabrid near acute tip. Culm 10-65 cm, erect or ascending, internodes with very short hairs below nodes and below panicle. Panicle 10-28 cm, very lax and nodding; branches spreading, slender, scabrid, tipped by 1-(2) spikelets. Spikelets 5.5-9.5 cm, 5-8-flowered, oblong-lanceolate to later wedge-shaped and gaping, greyish green to purplish. Glumes unequal, very narrow, acuminate, membranous, glabrous, but sparsely scabrid on keels and nerves in upper ½; lower 13-22 mm, 1-(3)-nerved, upper 20-30 mm, 3-(5)-nerved. Lemma 20-35 mm, 7-nerved, rounded, lanceolate, minutely scabrid, tapering to 2 acute to acuminate, hyaline, smooth to finely scabrid lobes 4-5 mm; awn 30-60 mm. Palea c. ⅔ length of lemma, keels sparsely ciliate. Callus with minute hairs. Rachilla 4-6 mm, finely, minutely scabrid. Stamens 2-3; anthers 1-3 mm. Caryopsis 11-13 × 1.5-2.8 mm. Plate 3A.
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Annual, caespitose. Culms 35–69 cm high. Leaves: basal sheaths puberulous to pilose (often sparsely so); blade 7–22 cm long, 2–6 mm wide, puberulous to pilose. Panicles nodding, open to contracted, 6.5–16 cm long; primary branches slightly spreading initially but becoming divaricate or reflexed. Spikelets spreading to pendulous, obovate, 27–35 mm long, with 6 bisexual florets; pedicel 5–60 mm long. Glumes subequal, apically keeled to dorsally rounded; lower glume 15.75–21 mm long, 1 (–3)-nerved with lateral nerves sometimes obscure, with midnerve apically papillose to scabrous and intercostal regions glabrous or sparsely puberulous; upper glume 22–31 mm long, 3 or 5-nerved, with midnerve apically scabrous and intercostal regions glabrous. Callus pointed, glabrous to scabrous or puberulous. Lemma 17–30 mm long, apically keeled, 7-nerved, scabrous; awn subapical, 36–60 mm long, straight, entered by 3 nerves. Palea slightly shorter than lemma, glabrous or minutely papillose or scabrous to sparsely puberulous. Anthers 0.75–3.5 mm long.
Annual; culms 35–80 cm. high, erect or spreading.. Leaf-blades 10–25 cm. long and 4–8 mm. wide; sheaths loosely hairy.. Panicles loose or somewhat contracted, up to 25 cm. long but variable in size and bearing few to many spikelets, nodding.. Spikelets loosely 5–8-flowered, oblong to wedge-shaped, 3–5 cm. long; glumes finely pointed, the lower subulate, 15–23 mm. long, 1–3-nerved, the upper narrowly lanceolate, 20–32 mm. long, 3–5-nerved; lemmas narrowly lanceolate in side view, 22–36 mm. long, herbaceous with hyaline margins, 7-nerved, scabrid, acuminately bilobed, awned from 4–7 mm. below the tip; awn 3.5–6 cm. long, straight, scabrid; callus oblong, ± 1 mm. long, obtuse; palea shorter than the lemma, sparsely and stiffly ciliate on the keels; anthers 2–3, 0.8–1.5 mm. long.
Loosely tufted annual, 0.3-1.1 m high; basal sheaths split. Leaf blade 50-400 x 3-8 mm, hairy; ligule an unfringed membrane. Inflorescence usually an open panicle; culm hairy below inflorescence; pedicels glabrous, scabrid or villous. Spikelet 30-90 x 3-8 mm (including awns), wedge-shaped, moderately laterally compressed; glumes unequal, shorter than spikelet; lower glume 1-nerved, upper 3-nerved. Florets 5-many; lemma 12-22 mm long, back rounded, apex acute, distinctly lobed; awn 30-70 mm long, stiff, straight; palea keels ciliate; ovary with a conspicuous hairy appendage at apex; anther 0.8-1.6 mm long. Flowering time Sept.-Jan., occasionally in Mar.
Loosely tufted annual 300-1100 mm high. Leaf blade 50-400 x 3-8 mm, hairy. Inflorescence usually open, branches stiff, erect or spreading; culm hairy below inflorescence; pedicels glabrous, scabrid or hairy. Spikelet 30-90 x 3-8 mm (including awns), open, wedge shaped, moderately laterally compressed; glumes unequal; lower glume 1-3-nerved, upper 3-5-nerved; lemma 12-22(35) mm long, rounded on back, apex distinctly bifid, acuminate; awn 30-70 mm long, stiff, straight, arising 1.5 mm or more below apex; palea keels ciliate; callus scar circular; anthers 0.8-1.6 mm long.
Annual; up to 1.1 m high; loosely tufted. Leaf blades 50-400 x 3-8 mm. Flowers: panicle usually lax and spreading; pedicels glabrous; scabrid or villous; spikelets 30-90 x 3-8 mm (including awns); spikelets open; wedge-shaped; lower glume 1-nerved; upper 3-nerved; lemmas 12-22 mm long; awns stiff and straight; 30-70 mm long; anthers 0.8-1.6 mm long.
Annual to 1 m. Leaves linear, hairy. Spikelets in a lax panicle, 30-90 mm long, lower glume 1-nerved, upper glume 3-nerved, lemma awn stiff and straight, 30-70 mm long.
A grass.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention -
Sexuality hermaphrodite
Pollination anemogamy
Spread epizoochory
Mature width (meter) -
Mature height (meter) 0.35 - 0.95
Root system -
Rooting depth (meter) 0.1
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Waste places and fallow land, also occurring in shallowly tilled crop land.
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A weed of disturbed sites andcrops. 
It is a temperate plant.
Light -
Soil humidity 1-3
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 5-11

Usage

Uses animal food
Edible seeds
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Bromus diandrus leaf picture by Jean-Marie NIER (cc-by-sa)
Bromus diandrus leaf picture by Kristell Riou-Nivert (cc-by-sa)
Bromus diandrus leaf picture by Ruth Puente (cc-by-sa)

Flower

Bromus diandrus flower picture by yochi (cc-by-sa)

Fruit

Bromus diandrus fruit picture by Bloom Blob (cc-by-sa)
Bromus diandrus fruit picture by Yoan MARTIN (cc-by-sa)
Bromus diandrus fruit picture by Yoan MARTIN (cc-by-sa)

Distribution

Bromus diandrus world distribution map, present in Australia, Austria, Canada, Colombia, Cyprus, Germany, Dominica, Algeria, Ecuador, Egypt, Spain, France, Georgia, Greece, Guatemala, Haiti, Indonesia, Iran (Islamic Republic of), Iraq, Italy, Jamaica, Kenya, Libya, Morocco, Malaysia, Namibia, Norfolk Island, Nicaragua, New Zealand, Palau, Portugal, Saudi Arabia, Sudan, Singapore, Turks and Caicos Islands, Tunisia, Tanzania, United Republic of, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:393675-1
WFO ID wfo-0000855460
COL ID NCXX
BDTFX ID 10663
INPN ID 86589
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Anisantha gussonei Anisantha macranthera Bromus boraei Bromus macrantherus Bromus gussonei Bromus diandrus f. glaber Bromus diandrus f. pilosus Bromus diandrus f. propendens Bromus murorum Bromus rigidus f. propendens Bromus propendens Anisantha diandra Bromus pilosus Bromus diandrus var. macrantherus Bromus gussonei var. boraei Bromus gussonei var. propendens Bromus maximus subsp. gussonei Bromus maximus subsp. macrantherus Bromus maximus var. glaber Bromus maximus var. gussonei Bromus maximus var. macrantherus Bromus maximus subsp. pilosus Bromus rigens subsp. macrantherus Bromus rigens var. gussonei Bromus rigidus subsp. gussonei Bromus rigidus var. gussonei Bromus rigidus var. macrantherus Bromus villosus subsp. gussonei Bromus villosus var. gussonei Bromus maximus var. pilosus Bromus diandrus f. boraei Bromus pallens Zerna gussonei Bromus diandrus