Annual or perennial herbs, sometimes woody at base, not armed. Lvs alternate, simple, entire or toothed. Capitula solitary or in loose corymbs. Involucral bracts in 1-2 rows, with membranous margins. Receptacle flat; scales 0. Outer florets ♀, ligulate; ligules yellow or orange. Inner florets tubular, with innermost functionally ♂ (some disc florets transformed into ligulate florets in cvs). Achenes heteromorphic, incurved, ridged, tuberculate or spiny on back; outer achenes narrowly beaked, or not beaked and 2-3-winged; inner achenes curled in ± 1 complete turn, usually without well-formed beaks or wings.