Plants small to medium-sized, in flattened mats, sordid pale green, yellowish, brownish, sometimes bluish, dull. Stems regularly or irregularly pinnate; hyalodermis absent. Leaves ± complanate, shriveled, often crisped when dry, ovate-oblong to lingulate, somewhat asymmetric, ventral leaves often more acuminate, dorsal and lateral leaves broadly oblong to oblong-ovate; margins plane, entire in basal 1/4, crenulate, serrulate, or strongly toothed in apical 1/4; apex rounded-truncate, short-apiculate; costa ending in apex, subpercurrent, divergent from base, parallel or somewhat convergent distally, distal abaxial surface toothed; laminal cells isodiametric or slightly longer than wide, irregularly hexagonal to quadrate, 1-papillose on both surfaces; marginal cells forming obscure border or not forming border. Specialized asexual reproduction unknown. Sexual condition synoicous and autoicous, infrequently dioicous; perichaetial leaf apex gradually acuminate to acute. Seta red to reddish yellow, moderately to strongly papillose-scabrous. Capsule inclined, horizontal, or pendulous, ovoid or oblong-cylindric; annulus absent; operculum abruptly short-rostrate or subulate. Calyptra mitrate-campanulate, base narrowed and shortly or deeply lobed, smooth or sometimes split on one side from base, somewhat scabrous or with few hairs distally.
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