Plants in small to medium-sized mats, white, yellowish, pale green, bluish, or brownish, pleurocarpous. Primary stems green, creeping, spreading, regularly or irregularly pinnate; hyalodermis present or absent, central cylinder cells large, walls thin, central strand absent; paraphyllia absent; pseudoparaphyllia absent [filamentous]; ± radiculose; axillary hairs of 2-several cells, basal cell small, brown, distal cell elongate, hyaline. Secondary stems spreading, suberect, frondose, not or sparingly branched. Leaves ± complanate, often dorsiventrally oriented, somewhat dimorphic; vertical (dorsal and ventral) leaves erect, broadly oblong, oblong-ovate, or ovate-lanceolate; lateral leaves erect-spreading to spreading, narrower; margins often dentate; apex usually acuminate; costa double, well developed; alar cells undifferentiated; laminal cells isodiametric or linear-rhomboidal, smooth to 1-papillose, walls lax to firm; marginal cells forming border or not. Sexual condition polygamous: autoicous, synoicous, or dioicous. Seta red to reddish yellow, elongate, smooth or papillose-scabrous. Capsule inclined, horizontal, pendulous, or sometimes erect, ovoid to oblong-cylindric, smooth, contracted below mouth when dry; annulus absent or narrow; operculum conic, rostrate to subulate; peristome double, dark red or red-brown; exostome teeth 16, lanceolate to lance-subulate, furrowed, cross striolate basally, papillose distally, trabeculate, sometimes bordered; endostome yellow to yellow-brown, finely papillose to nearly smooth, basal membrane high, segments lanceolate, as long as teeth, keeled, cilia absent. Calyptra conic-mitrate to mitrate-campanulate, base lobed or fringed, smooth or papillose-scabrous distally, naked. Spores smooth to finely papillose.
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