Petals unguiculate (sometimes very shortly), sometimes auriculate or hastate at the base, glabrous, the margins wholly or partially fimbriate.
Fruit with lateral wing completely surrounding the nut, circular or broadly elliptic; dorsal wing small, obliquely lanceolate or absent.
Leaves spirally arranged or opposite, with 2 glands on the margin near the base or eglandular; stipules very small, deciduous.
Woody climbers or upright or semiscandent shrubs; stems with younger parts usually ± densely appressed-pubescent or sericeous.
Ovary densely sericeous; styles truncate, shorter than, as long as, or slightly exceeding the stamens.
Inflorescences racemose or corymbose-paniculate, axillary or terminal.
Stamens glabrous with dorsifixed anthers.
Flowers actinomorphic, bisexual.
Sepals without glands.