Celtis philippensis Blanco

Species

Angiosperms > Rosales > Cannabaceae > Celtis

Characteristics

Small to large tree, up to 30 m, 80 cm ø. Buttresses if present up to 2½ m tall, 2 m wide and 10 cm thick. Bark smooth to finely fissured, pale grey to grey-brown. Innovations initially sparsely to densely set with yellowish-brown appressed or/and woolly hairs, glabrescent. Stipules ovate-acute, 6-10 by 2-4 mm, thick, peltately attached, overlapping and enclosing the bud. Leaves thick-coriaceous, glabrous, full grey when dried, elliptic-oblong to suborbicular, (4-)8-14(-18) by (2-) 3-6(-8)cm, index (l½-)2-3; base rounded or attenuate-rounded, mostly symmetrical; margin entire often undulate (immature leaves very rarely distantly serrulate at the upper half); apex rounded to acute; midrib and nerves raised beneath, impressed to flattish above; main nerves 1 pair, ascending, arcuating and running throughout the length of the leaf (var. philippensis) or 1-3 pairs, the lowest pair ascending, arcuating, and running up to about ⅔ the length of the leaf and then anastomosing with the 1-2 weaker and more or less horizontal upper nerves (var. wightii); reticulations fine, dense, subscalariform or subareolate, usually rather distinct beneath; petiole 6-15 by 1-2 mm, sulcate. Inflorescences ♂ or ♂ ⚥, much-branched panicles, many-flowered, including the bracts densely yellow-brown to rufous soft-hairy; bracts ovate-acute, c. 3 by 1 mm. In the ♂ ⚥ inflorescence the ⚥ flowers are borne on the distal parts of the inflorescence. ♂ Inflorescences borne on the lower part of the new shoots, 2-4 cm long, with up to 40 flowers; ♂ flowers c.   ø; perianth lobes ovate-elliptic, c. 1½-2 by 1 mm; filaments 1-1½ mm long, anthers subreniform, c. ½-1 mm by ½ mm; pistillode ovoid-cylindrical, compressed, c. 1-1½ by ½ mm. — Mixed inflorescence up to 5 cm long, up to 50-flowered, borne on the upper part of the new shoots; ⚥ flowers ovoid, c. 2-2½ by 2 mm; perianth lobes ovate-elliptic, c. 2-2½ by 1 mm; filaments 1-2 mm, anthers ½-l mm ø; ovary ovoid-cylindrical, c. 2-2½ by 1½-2 mm, glabrous except at the base; stigmatic arms spreading, c. 1-  long, bilobed to bifid at the tip. Infructescence up to 4-5 cm long, carrying 1-3 fruits, axes 1-2 mm thick. Fruit ovoid, glabrous, 8-15 by 7-12 mm, beaked when young; exocarp less than 1 mm ø, sometimes lenticellate, turning orange to red when ripe; endocarp ± smooth; embryo curved, hypo-cotyle ascending, cotyledons broad, foliaceous, unequal in thickness, not folded; endosperm oily, scanty to absent.
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Trees to 30 m tall, evergreen. Bark grayish white to grayish brown or gray. Branchlets brown or tan, glabrous or pubescent. Winter buds naked, without differentiated scales. Stipules ovate-lanceolate to ovate, 2-8 mm, glabrous or pubescent, enfolding naked terminal bud, apex attenuate. Petiole glabrous, 0.5-1.5(-2) cm, adaxial furrow deep and narrow or broad and shallow; leaf blade lanceolate-elliptical, ovate-elliptical, or oblong, 4.5-18 × 2.5-11.5 cm, papery to leathery, glabrous or with a scattering of inconspicuous appressed hairs, base acute, obtuse, or rounded, symmetric or weakly asymmetric, margin entire, apex rounded-obtuse to gradually or abruptly acuminate; secondary veins 1 or 2 on each side of midvein. Cymes 1 or 2 per leaf axil, densely flowered, much branched, each with ca. 5 bisexual flowers and 5 or more male flowers. Styles 2, apically broadened and 2-cleft or emarginate. Infructescences 1-3 per leaf axil, stout, branched or unbranched, with 1-3 drupes and prominent scars from fallen flowers, glabrous, 2.5-5 cm, 3-6 × as long as subtending petiole. Drupe orange to red, 0.8-1.4 cm, globose to ellipsoid, base rounded to obtuse, apex rounded to obtuse. Stone ovoid to ± globose, 6-13 mm, conspicuously to inconspicuously ribbed.
Shrub or tree to 15 (–25) m high. Adult leaves: lamina elliptic to suborbicular or ovate to broadly ovate, (1.3–) 4–21 cm long, 0.9–10 cm wide, rounded at base, sometimes oblique, mostly entire (sometimes toothed), acuminate, markedly 3-veined, glabrous to slightly scabrous above, glabrous to sparsely puberulous below; petiole 2–14 mm long; stipules ovate, 6–10 mm long, peltate, caducous. Inflorescence a many-flowered panicle, either male or male and bisexual. Flowers 5-partite. Male flowers: perianth lobes ovate-elliptic, 1.5–2 mm long; stamens 1.5–2.5 mm long; anthers subreniform. Bisexual flowers: perianth lobes ovate-elliptic, 2–2.5 mm long; stamens 1.5–3 mm long; ovary ovoid-cylindrical, 2–2.5 mm long; stigmatic arms 1–1.5 mm long, bilobed or bifid. Drupe ovoid, 7–15 mm long, yellow, orange, pink or red; pedicel 0.3–1.5 cm long.  See also Dixon & Kerrigan (2011), Zich et al. (2020). [The above description of Celtis philippensis is adapted from Hewson (1989) but may require further revision to exclude elements of the species Celtis strychnoides which was included as a synonym of C. philippensis at the time of publishing the Flora of Australia Volume 3 treatment-Editor, 23 September 2021.]
A medium sized tree. It grows 4-10 m tall. It often has buttresses. It has a dense rounded crown. The tree can lose many of its leaves during the dry season of the year. The bark is smooth with a mottled cream grey pattern on it. The leaves are alternate and smooth. They are stiff and brittle. They are oval and about 3-12 cm long by 2-6 cm wide. The leaves are dark green above and paler underneath. There are 3 easy to see veins along the leaf. These are raised and white, underneath the leaf. The leaves have a pointed tip and short stalk (1 cm). The young leaves have teeth along the edge. The flowers are small and white and 2-3 mm across. They occur in small clusters (2 cm across) in the axils of leaves. Flowers of one or both sexes can occur on the same plant. The fruit are smooth and fleshy. They are round but flattened. They can be 0.7-1 cm across. They are yellow, but bright red when ripe. They contain a single hard shelled seed. The seed can be 6 mm across.
Leaves (5)6.5–12(17.5) x 2.5–4.5(8) cm., elliptic to elliptic-ovate or elliptic-oblong; apex with a wide (usually short) acumen and mucronate tip; base slightly asymmetrical and broadly cuneate to rounded or subcordate; margin entire or (rarely in Flora Zambesiaca region) coarsely dentate in the upper half; lamina thinly coriaceous, pale green, punctate but smooth and ± shiny above, glabrous; 3-nerved from the base, the basal lateral nerves extending almost to the apex, as strongly prominent beneath as the midrib, upper lateral nerves 1–2(3) on each side of the midrib, rather fine and inconspicuous; secondary venation fairly closely parallel, ± horizontal between the midrib and basal lateral nerves.
Inflorescence in leaf axils towards the ends of branches; lower inflorescences 5–30 mm. long, containing many crowded male (lowers, these sessile or with pedicels to 2 mm. long, and a few female and bisexual flowers at apices of branches of these inflorescences, their pedicels usually longer; upper inflorescences usually short with several bisexual flowers.
Evergreen much-branched tree 5–20 m. tall, monoecious; bole often with short buttresses, bark smooth grey, wood white; young stems and branches very sparsely white-pubescent, glabrescent.
Petiole 4–10(16) mm. long; stipules 3–7 mm. long, lanceolate, shortly produced below the point of attachment, ± pubescent.
Ovary ± glabrous, with a basal ring of long hairs; styles very shortly bifid, (1)1.5–2(3.5) mm. long.
fruits red, 9–12 x 7–10 mm., ovoid, glabrous; pedicels 3–7 mm. long.
Sepals 1.5–2.5 mm. long, pubescent.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention evergreen
Sexuality hermaphrodite
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Mature height (meter) 5.0 - 20.0
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Nitrogen fixer -
Photosynthetic pathway c3

Environment

Grows in a wide range of habitats and in a wide range of soil types, including basalt, quartzite, sandstone, limestone, coastal sand and black peaty soil. Habitats include open sclerophyll forest, rainforest, monsoon forest, vine forest and coastal dunes.
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It is a tropical plant. It grows in coastal monsoon forests. It can be on sand dunes and is often near permanent water. It grows up to 1,200 m above sea level. It needs well drained soil. It suits hardiness zones 10-12. In XTBG Yunnan.
A canopy or subcanopy tree in lowland, groundwater and riverine forest. Semideciduous rain-forests; gallery forest; swamp forest; forest regrowth; savannah near forest; often locally abundant; at elevations from 229-1,320 metres.
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Hardiness (USDA) 9-12

Usage

The fruit are eaten raw when ripe. The seed oil is edible.
Uses environmental use fiber fuel invertebrate food material medicinal oil poison social use wood
Edible fruits leaves seeds
Therapeutic use -
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Animal toxicity -

Cultivation

Plants can be grown from fresh seed.
Mode seedlings
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Images

Fruit

Celtis philippensis fruit picture by Noel Dionson (cc-by-sa)

Distribution

Celtis philippensis world distribution map, present in Angola, Australia, Benin, Brunei Darussalam, Central African Republic, China, Côte d'Ivoire, Cameroon, Congo, Ethiopia, Gabon, Ghana, Guinea, Equatorial Guinea, Hong Kong, Indonesia, India, British Indian Ocean Territory, Kenya, Liberia, Sri Lanka, Madagascar, Myanmar, Mozambique, Mauritius, Malaysia, Mayotte, Nigeria, Philippines, Papua New Guinea, Réunion, Senegal, Solomon Islands, South Sudan, Sao Tome and Principe, Togo, Thailand, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Viet Nam, and South Africa

Conservation status

Celtis philippensis threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:56400-3
WFO ID wfo-0000593688
COL ID 5XGJQ
BDTFX ID -
INPN ID 807238
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Sponia strychnifolia Celtis brevinervis Celtis collinsae Celtis djungiel Celtis multifolia Celtis trinervia Celtis wightii Solenostigma consimile Celtis laurifolia Celtis philippensis Celtis hasseltii Celtis mindanaensis Solenostigma brevinerve Bosea trinervia Celtis insularis Celtis philippensis var. wightii Celtis philippensis var. consimilis Celtis wightii var. consimilis Celtis philippensis var. philippensis