Celtis L.

Micocoulier (fr)

Genus

Angiosperms > Rosales > Cannabaceae

Characteristics

Small to large monoecious or polygamo-monoecious trees, often buttressed. Bark smooth or finely fissured, often conspicuously warty lenticellate. Branches (in Mal.) unarmed, initially densely yellow-brown or rufous-hairy, glabrescent; hairs 1-celled. Buds enclosed by the overlapping stipules or naked. Stipules thick and tough, peltately attached or free and scarious, caducous. Leaves entire 3r not, 3-nerved at base, semideciduous or persistent. Inflorescenses ♂, ⚥, or ♂⚥, branched racemes or panicles, few-to many-flowered, axillary or subterminal on the new shoot; staminate inflorescences borne on the lower and leafless part or in the axil of leaves of the new shoot; in the ♂⚥ inflorescence the ⚥ flowers are borne on the distal ends of the axes; bracts minute, caducous. — Staminate (♂) flowers globular, pedicelled or sessile; perianth lobes 4-5, imbricate in bud, membraneous, boat-shaped, outside sparsely pubescent, at anthesis recurved, caducous; stamens glabrous, inserted on the densely pilose receptacle; filaments subulate, incurved in bud and spreading elastically, exserted at anthesis; anthers ovoid to subreniform, dorsifixed just above the emarginate base, extrorse; pistillode present or absent. ⚥ Flowers ovoid, pedicelled; perianth lobes 4-5, imbricate in bud, connate at base, membranous, outside sparsely pubescent, boat-shaped, at anthesis recurved, caducous; stamens well-developed and functional or rudimentary, other characters as in ♂ flowers; ovary ovoid-ellipsoid, sessile, style short or ± absent; stigmatic arms elongate, divergent, the tips entire to deeply bifid; ovule anatropous. Drupe fleshy, ovoid, ellipsoid or globose; exocarp thick and firm, mesocarp thin and fleshy, containing slimy substances; endocarp hard and persistent, smooth or variously ridged or pitted. Seed: coat membranous, chalazal area broad, dark-coloured and close to the minute hilum; endosperm scanty or wanting, oily or gelatinous, nearly enclosed between the folds of the cotyledons. Embryo curved, cotyledons broad, foliaceous, equal or unequal in thickness, flat or conduplicate, variously folded, incumbent on or embracing the short superior and ascending radicle. Germination epigeal.
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Polygamo-monoecious or monoecious shrubs or trees, unarmed or with spinose branches. Leaves alternate, distichous, often inequilateral, serrate or entire; peti-olate; stipules lateral. Inflorescences borne on this year's wood: staminate inflorescence cymose or fasciculate, pistillate inflorescence solitary or few-flowered fasciculate. Flowers small, pedicellate. Bisexual flowers: sepals 4-5, more or less connate at the base; stamens as many as and opposite the calyx lobes, the anther dehiscence extrorse, disc present; ovary sessile, 1-locular, the styles 2, sometimes united at the base, reflexed. Staminate flowers with rudimentary pistil. Pistillate flowers without staminodes. Fruit an ovoid or subglobose drupe, the embryo curved, the cotyledons broad, conduplicate or rarely flat, variously folded.
Trees or shrubs, evergreen or semideciduous, monoecious, sometimes with some bisexual flowers. Leaves alternate, 3-veined at base; stipules lateral, free, scarious, enclosing bud, caducous. Inflorescence axillary on new shoots, a raceme or panicle; bracts minute, caducous. Male flowers globular; perianth lobes 4 or 5, imbricate in bud, free, recurved at anthesis; stamens 4 or 5, exserted at anthesis; anthers dorsifixed, extrorse; pistillode present or absent. Bisexual flowers ovoid; perianth lobes 4 or 5, as for male flowers; stamens 4 or 5; ovary 1-locular, sessile; style short or absent; stigmas elongate, entire or bifid, divergent; ovule pendulous. Female flowers: as for bisexual flowers but stamens reduced to staminodes. Fruit a drupe, ovoid or globose. Seed without endosperm; embryo curved.
Trees or shrubs, monoecious or rarely dioecious, ± armed. Leaf-blades penninerved or 3–5-nerved from the base, which is often unequal-sided, and with cystoliths, giving a scabrous texture to the mature leaves, often present. Stipules usually small and caducous. Inflorescences cymose or fasciculate, the lower ones usually with many predominantly ♂ flowers, the upper ones with few longer pedicelled ♀, hermaphroditeflowers. Sepals (4–)5, free or almost so, imbricate. Stamens as many as the sepals; anthers ovate. Ovary sessile, 1-locular; styles ± 1–2-branched. Drupes thinly fleshy, subglobose to ellipsoid; endocarp hard, sometimes ribbed.
Leaves petiolate, lamina usually unequal-sided at the base, penninerved or 3-nerved from the base; cystoliths often present, giving a scabrous texture to the mature leaves; stipules usually small.
Climbers, shrubs or treelets with spines, monoecious. Inflorescences bisexual or unisexual. Tepals 5; stamens 5 and opposite the tepals; stigmas furcate. Fruit at least 10 mm long.
Flowers often precocious, in many small cymes (sometimes fasciculate in non African species), female flowers sometimes in separate inflorescences.
Trees or shrubs, monoecious rarely dioecious, unarmed (often armed in non African species).
Fruit drupaceous, thinly fleshy, sometimes with ribbed endocarp.
Ovary sessile; styles often once or twice-branched.
Sepals (4)5, ± free, imbricate.
Anthers ovate.
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Foliage retention evergreen
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Environment

Malesian species of Celtis may be classified into two rather distinct ecological groups, l.c. the C. philippensis var. philippensis and C. tetrandra groups. The first group, which includes also C. hilde-brandii, C. latifolia, C. luzonica, C. paniculata, and C. rigescens, is found mainly in the lowland forests, both primary and secondary, and is an important constituent of the understorey tree community in moist areas. The C. tetrandra group, which includes C. rubrovenia, C. timorensis, and C. philippensis var. wightii, is confined to areas which are subject to a rather pronounced seasonal climate, or if they occur in wetter regions, they grow on strongly drained substrates, e.g. rocky shores, limestone, etc. In tune with this environmental preference, the second group shows a more prominent flush-wise growth habit and its species are completely or partly deciduous. In Malaya, species of Celtis are producing flowers around July-September, while fruit ripens January-March.How pollination is affected is not certain, but judging from the structure and position of the inflorescence, some species (e.g. C. philippensis var. philippensis, C. hildebrandii, and C. latifolia) may be pollinated by insects, while others (C. tetrandra, C. timorensis, C. rubrovenia, and C. rigescens) may be pollinated by wind.The ripe fleshy drupes which turn to orange, red or bluish-black may be dispersed by birds, or alternatively they may be dispersed by water as the embryo is protected by the hard, persistent and durable endocarp.
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Hardiness (USDA) 4-10

Usage

Widely cultivated. See Rodd (1996), Spencer (1997).
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Cultivation

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