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Inflorescences cymose, often branched, usually congested, entirely male or with 1(2) females near the base; remaining female flowers solitary and usually in the upper axils.
Leaves penninerved, shortly petiolate, apex long-mucronate, base slighty unequal-sided; stipules large, united along one margin, enclosing the terminal bud, caducous.
Sepals 5, shortly basally united, with the male buds induplicate-valvate, and the female buds imbricate.
Female flowers without staminodes; male flowers with pistillode, stamens equal in number to sepals.
Ovary sessile, unilocular; styles 2, long, unbranched, divaricate, persistent.
Trees or shrubs, monoecious or dioecious; branches with axillary spines.
Fruits large, endocarp very hard.
Embryo curved.
