Leaves opposite or in whorls of 3–4, mostly drying pale, petiolate or subsessile, usually deciduous; nodules absent; domatia small, white-pubescent or absent; stipules mostly short, ovate or triangular, entire, bifid or sometimes with a few teeth.
Flowers small, heterostylous, (4)5(6)-merous, hermaphrodite, in mostly small, sessile or pedunculate inflorescences, often in small heads, or sometimes paniculate, ± sessile or pedicellate; bracts and bracteoles very small or absent.
Fruit a drupe with 2 pyrenes; pyrenes ± flat ventrally, mostly ± 3-ribbed or 3-lobed dorsally, opening by 2 slits extending along the margins of the ventral face for about half its length.
Shrubs, often flowering before the leaves are fully developed; stems usually 2-ribbed, all but the youngest shoots usually covered with pale brown soft cork.
Stamens with long filaments in short-styled flowers and shorter filaments in long-styled flowers, the anthers included or only the tips exserted.
Ovary 2-locular, each locule containing a single erect ovule; style filiform, usually papillate-pubescent, divided into 2 thick stigma lobes.
Corolla mostly yellow or white; tube shortly cylindrical, hairy at the throat; lobes triangular to elliptic-lanceolate, always valvate.
Calyx limb usually short, truncate or toothed.
Seeds pale; endosperm not ruminate.