Shrubs or trees, evergreen, perennial, monoecious; stems and foliage without latex. Indumentum of simple, multicellular trichomes, glandular trichomes absent, stinging trichomes absent. Stipules entire, inconspicuous, deciduous. Leaves opposite, petiolate, elobate, penniveined, entire or crenate to serrate, glands absent. Inflorescences axillary, racemose, solitary or fascicled, usually unisexual, with flowers in bracteate clusters. Male flowers pedicellate; calyx lobes 2+2 or 3+3, imbricate, free and ± equal; petals absent; disc absent; stamens 4 or 6, filaments free and attached to a slightly raised receptacle; anthers dorsifixed, bilobate, thecae globose and longitudinally dehiscent; pistillodes present. Female flowers pedicellate; calyx lobes 3+3, imbricate, free; petals absent; disc absent; ovary 3-or 4-locular, ovules biloculate; styles shortly connate at base, simple. Fruit capsular, trilobate, surface smooth, dehiscing septicidally into 3 bivalved cocci. Seeds ovoid; testa crustaceous; albumen fleshy; caruncles entire, non-arilloid; cotyledons broad, flat.