Cleomaceae Bercht. & J.Presl

Cléomacées (fr)

Family

Angiosperms > Brassicales

Characteristics

Herbs or shrubs, annual or perennial (usually deciduous, evergreen in Peritoma arborea); spines usually absent (present in Hemiscola and Tarenaya); glabrous or glandular-pubescent, hairs stalked or sessile (producing glucosinolates). Stems usually erect, sometimes spreading or procumbent; branched or unbranched. Leaves alternate, spirally arranged (usually palmately compound, sometimes simple); venation pinnate; stipules usually present (usually caducous, sometimes deciduous, 3-8-palmatifid, linear, threadlike, minute, scalelike, or absent, nodal (stipular) spines present in Tarenaya and Hemiscola); petiole present (pulvinus usually present, nectaries absent, petiolar spines sometimes present, petiolules present); blade margins entire, serrate, or serrulate. Inflorescences terminal or axillary, usually racemose, sometimes flat-topped, or flowers solitary (usually elongated in fruit); bud scales absent; bracts present or absent (unifoliate, often trifoliate proximally, bracteoles absent). Pedicels present. Flowers usually bisexual (developmentally unisexual within sections of racemes), actinomorphic or slightly zygomorphic, rotate to crateriform, campanulate, or urceolate; perianth and androecium hypogynous; sepals persistent or deciduous, 4, distinct or connate basally; petals 4, attached directly to receptacle, imbricate, distinct, equal or unequal; intrastaminal nectary-discs, scales, or glands present or absent; stamens [4-]6-27[-35]; filaments free or basally adnate to gynophore (or along proximal 1/3-1/2 in Gynandropsis) or androgynophore, glabrous or pubescent; anthers dehiscing by longitudinal slits, pollen shed in single grains, binucleate, commonly tricolporate; gynophore present or absent; pistil 1; ovary 1-carpellate (except 2 in Oxystylis), 2-locular; placentation parietal; ovules 1-18(-26+) per locule, anatropous, bitegmic; style 1 (straight, relatively short, thick, not spinelike in fruit, except in Oxystylis, sometimes in Wislizenia); stigma 1, capitate, unlobed. Fruits capsular or nutlets (usually stipitate from elongation of gynophore, erect to divergent, usually not inflated), valvate, elongate (± dehiscent by 2 lateral valves, except in Polanisia), or schizocarps (inflated in Peritoma arborea), indehiscent or dehiscent. Seeds 1-65[-200], tan, yellowish brown, light brown, pale green, brown, reddish brown, silver-gray, or gray to black (papillose or tuberculate); arillate or not; endosperm scanty or absent, persistent perisperm sometimes present; cotyledons incumbent, (radicle-hypocotyl elongated).
More
Herbs [rarely shrubs], sometimes woody at base, producing mustard oils. Stems erect, sparsely or profusely branched, glabrous or glandular pubescent. Stipules scalelike or absent, caducous (petiolar spines sometimes present). Leaves alternate, spirally arranged, palmately compound; petiole often pulvinate; leaflets [1 or]3-7[-11]; leaflet blades with pinnate venation. Inflorescences racemes or corymbs or flowers solitary and axillary; peduncle present; bract present [or absent] at base of pedicels. Pedicel present; bracteoles absent. Flowers bisexual but sometimes appearing unisexual due to incomplete development, actinomorphic or slightly zygomorphic, rotate, crateriform, campanulate, or urceolate, hypogynous. Sepals 4, distinct or basally connate, persistent. Petals 4, distinct, imbricate, attached directly to receptacle; intrastaminal nectary-disk or glands present or sometimes absent. Stamens 6(-32); filaments free or basally adnate to gynophore (or along basal 1/3-1/2 in Gynandropsis); anthers dehiscing by longitudinal slits; pollen shed in single grains, 2-nucleate, commonly 3-colporate. Pistil 1, 2-carpellate; ovary superior; ovules 1 to many per locule, 2-tegmic, anatropous, placentation parietal; style 1, straight, short, thick; stigma 1, capitate, unlobed. Fruit an elongate capsule, ± dehiscent by lateral valves along their entire length [indehiscent or dehiscent schizocarp], usually stipitate from elongation of gynophore (lacking in Arivela). Seeds 1-10(-40) per capsule, tan, yellowish brown, or brown, cochleate-reniform, papillose or tuberculate, arillate or not; endosperm scanty or none but a persistent perisperm sometimes present.
Annual or perennial herbs or subshrubs, rarely shrubs; glabrous or indumentum of simple glandular or eglandular hairs; stems sometimes armed. Leaves alternate, usually palmately compound, sometimes simple; stipules usually present; leaflets usually 3–7; venation pinnate. Inflorescence axillary or terminal, a raceme or corymb, sometimes flowers solitary, usually bracteate. Flowers bisexual, actinomorphic to slightly zygomorphic. Sepals 4, free or fused basally, sometimes deciduous; petals 4, free, equal or unequal, variously coloured. Stamens 6–27 (–35), sometimes some or all reduced to staminodes; filaments free or basally fused to gynophore, nectaries sometimes associated with stamens; anthers dehiscing by longitudinal slits. Gynoecium usually 1-or 2-carpellate. Ovary superior, 2-locular, ovules 1–many per locule; style 1, stigma capitate; placentation parietal. Fruit usually a dry pedicellate capsule, usually dry lateral valves, with a persistent, thin, hoop-like woody placenta. Seeds reinform or horseshoe-shaped to coiled or conduplicate, smooth to tuberculate, often exarilate.
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Sexuality hermaphrodite
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Usage

Some species of Cleome (Spider Flowers) are grown as ornamentals.
Uses ornamental
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Cultivation

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Images

Cleomaceae unspecified picture

Distribution

Cleomaceae world distribution map, present in Australia and China

Identifiers

LSID -
WFO ID wfo-7000000137
COL ID 8BN
BDTFX ID 101122
INPN ID 452731
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Cleomaceae

Lower taxons

Cleomella Dactylaena Cleome