Cleome L.

Carotte intermédiaire au sens large (fr)

Genus

Angiosperms > Brassicales > Cleomaceae

Characteristics

Herbs, annual, sparsely branched, glandular pubescent or glabrous, lacking spines. Stipules absent or scalelike and caducous. Leaves alternate, spirally arranged, pinnately compound; petiole long or short, with a pulvinus at basal or distal end; leaflets [1 or]3-7[-11]; petiolule bases connate, forming a pulvinar disk; leaflet blades lanceolate to oblanceolate, margin entire or serrulate. Inflorescences terminal or axillary from apical leaves, racemose, sometimes flat-topped or elongated, elongating in fruit; bract present [or absent] at base of pedicels. Flowers slightly zygomorphic. Sepals 4, equal, connate for ca. 1/2 their length, each often subtending a basal nectary. Petals 4, distinct, equal. Stamens [4-]6, distinct; filaments inserted on a discoid or conical receptacle (androgynophore). Gynophore slender, elongating and recurving in fruit or sometimes obsolete; carpel 1; style 0.2-0.8 mm, thick; stigma 1, capitate. Fruit an oblong capsule, dehiscent. Seeds 10-40 per capsule, reniform, arillate or not; cleft fused between 2 ends of seed.
More
Annual (or perennial) herbs, often hairy, sometimes glandular-hairy. Stipules none or obsolete; seldom with stiff, recurved, thorn-like, stipular (?) enations at the base of the leaf. Leaves petioled, herbaceous, in the Mal. spp. palmately dissected into 3-7 leaflets. Flowers pedicelled, in leafy, terminal racemes or panicles, the leaves apically gradually reduced, mostly slightly zygomorphic in the position of the petals. Sepals 4, free. Petals (normally) 4, the base often clawed. Stamens 6 to ∞, in Mal. spp. all fertile, sometimes at the base connate with the gynophore to an androgynophore. Disk small. Ovary 1-celled, sometimes sessile but mostly on a gynophore; stigma knob-shaped or flattish, subsessile. Capsule linear, terete, 2-valved, beaked, dehiscing from the base or from the apex, the 2 placentae forming a persistent replum. Seeds ∞, orbicular to horseshoe-shaped (with a more or less open cleft), sometimes with a funicular elaiosome, on the dorsal side sculptured to scaly.
Herbs, sometimes suffrutescent, rarely clambering, frequently glandular-pubescent or thorny; leaves alternate, stipulate or exstipulate, usually palmately compound, occasionally simple, the leaflets entire or minutely callose-serrulate; inflorescence racemose, terminal or both terminal and lateral, few-to many-flowered, bracteate, rarely ebracteate; calyx deeply 4-parted, persistent or decidu-ous; petals 4, more or less unequal, usually unguiculate; disc usually present, symmetrical or asymmetrical, rarely absent; stamens 6, rarely 4, inserted on a short or more or less elongate androgynophore, occasionally nearly sessile, the fila-*ments more or less unequal and declinate; ovary borne upon a more or less elongate gynophore, rarely nearly sessile, the stigma sessile or stipitate, the ovules numerous; fruit a dry, terete silique, dehiscing from the persistently joined replum by 2 valves; seeds cochleate-reniform, smooth, minutely tuberculate, or transversely rugose.
Herbs, annual or perennial. Stems unbranched or sparsely branched ; glandular-pubescent, glabrous, glabrescent, or scabrous. Leaves: stipules absent or scalelike; petiole with pulvinus basally or distally, (petiolule basally adnate, forming pulvinar disc); leaflets 1 or 3[-11] (flat). Inflorescences terminal or axillary (from distal leaves), racemes (flat-topped or elongated); bracts present [absent]. Flowers zygomorphic; sepals persistent, basally connate (1/2 of length), equal (each often subtending a nectary); petals equal; stamens [4] 6; filaments inserted on a discoid or conical androgynophore, glabrous; anthers (oblong to linear), coiling as pollen is released; gynophore recurved in fruit [obsolete]. Fruits capsules, dehiscent, oblong. Seeds 4-25, reniform or ovoid-spheroidal, arillate or not, (cleft fused between ends). x = 10 (?).
Herbs, mainly annual in East Africa, sometimes bushy and woody below. Leaves alternate, petiolate, simple or digitately compound with 3–5–7–9 very shortly petiolulate leaflets. Inflorescence racemose; flowers pedicellate, subtended by foliaceous or membranous, persistent or caducous bracts. Sepals 4, free or shortly united at the base. Petals 4, equal or unequal, often long-clawed. Androphore absent; stamens free; filaments equal or unequal, sometimes medially swollen; staminodes present or absent. Gynophore present or absent; ovary 1-locular with many parietal ovules. Fruit a siliquiform capsule with 2 dehiscing valves and a persistent replum. Seeds discoid or reniform, the ends incurving or one of them free; testa smooth, rugulose or ridged.
Pet equal or nearly so, entire; receptacle bearing an adaxial, usually rounded or pointed, mostly greenish or yellowish gland between the cor and the androecium; style mostly (incl our spp.) very short; frs usually long-stipitate, reticulately veined, opening from the base upwards, the valves deciduous; annuals with palmately compound or trifoliolate lvs, simple bracts, and handsome fls in elongating terminal racemes. 200, mainly warm reg.In addition to the following spp., the Middle-Eastern C. ornithopodioides L. is adventive on chrome ore piles in Ky. It is slender, to 5 dm, with strongly glandular stem, trifoliolate lvs (the lfls to 1.5 cm) and small fls (the pet only 1–2 mm) and frs (1.5–3.5 cm).
Annual or perennial herbs, glabrous or glandular-pubescent. Stipules usually absent. Leaves palmately dissected, of 3–7 leaflets. Flowers solitary-scapose or in terminal raceme. Sepals free, equal. Petals with or without claw, equal, often oriented to adaxial side. Stamens 1–30, sometimes connate with each other at base and adnate to gynophore to form androgynophore. Receptacular gland variable from concentric to unilateral. Gynophore present or absent; ovary unilocular with 2 placentas; stigma capitate. Fruit a capsule. Seeds many, orbicular to horseshoe-shaped, sculptured, with or without an aril.
Herbs, sometimes subshrubs. Lvs simple or digitate-compound. Fls solitary or in racemes, actinomorphic or subzygomorphic. Sepals 4, fused or free, persistent or caducous. Petals 4, sessile or clawed, equal or subequal. Stamens (4)-6-(10), free or adnate at base to form a short androgynophore; anthers usually exserted. Ovary on a gynophore; stigma sessile or style very short. Fr. a siliquiform capsule, glabrous or hairy. Seeds many, small, reniform.
Fruit an oblong or linear capsule often borne on an elongated gynophore, with two valves separating from the seed-bearing placentas, often tipped with the persistent style; valves glabrous or pubescent, smooth or strongly longitudinally nerved.
Stamens 2 — many, all fertile or some sterile, usually borne on a small torus or receptacle; filaments equal or unequal and declinate, occasionally thickened at the apex; anthers 2-thecous, usually oblong or narrowly oblong.
Ovary sessile or with a short gynophore, with many ovules on two parietal placentas; style short or absent; stigma capitate or truncate.
Leaves usually petiolate, simple or digitately 3–9-foliolate; leaflets usually narrow, linear to lanceolate, entire or serrulate.
Seeds reniform, smooth or ridged transversely, radially or longitudinally, or reticulate-tuberculate, glabrous or pubescent.
Sepals 4, usually free or almost so, usually narrow and often glandular or setulose-pubescent.
Annual or perennial herbs or occasionally small shrubs.
Petals 4, sessile or clawed, equal or unequal.
Flowers ± zygomorphic, in terminal racemes.
Life form annual
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

In Malaysia most species are weeds along roadsides and in fields at low altitudes.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 4-10

Usage

Uses. Two spp. are cultivated as ornamentals, especially C. speciosa. Some are used as vegetables and in primitive medicine.
Uses medicinal ornamental
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 10
Germination temperacture (C°) 21 - 23
Germination luminosity light
Germination treatment stratification
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Cleome unspecified picture

Distribution

Cleome world distribution map, present in American Samoa, Australia, China, Malaysia, New Zealand, Panama, Somalia, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:30009982-2
WFO ID wfo-4000008580
COL ID 62L7Q
BDTFX ID 101964
INPN ID 612457
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Justago Polanisia Tarenaya Buhsia Isexina Podandrogyne Puccionia Haptocarpum Cleoserrata Physostemon Dipterygium Cleome Gynandropsis Chilocalyx Cristatella Pedicellaria Coalisina

Lower taxons

Cleome africana Cleome aldenella Cleome amblyocarpa Cleome anomala Cleome arabica Cleome brachystyla Cleome brasiliensis Cleome burttii Cleome carnosa Cleome coeruleo-rosea Cleome costaricensis Cleome decipiens Cleome densifolia Cleome dichotoma Cleome diffusa Cleome dodecandra Cleome dodecaphylla Cleome elegantissima Cleome eosina Cleome felina Cleome fimbriata Cleome foliolosa Cleome foliosa Cleome fosteriana Cleome glandulosa Cleome glaucescens Cleome gobica Cleome gossweileri Cleome houstonii Cleome houtteana Cleome humilis Cleome iberica Cleome kenneallyi Cleome kermesina Cleome kersiana Cleome khorassanica Cleome laburnifolia Cleome lanceolata Cleome latifolia Cleome longipes Cleome macradenia Cleome macrantha Cleome macrophylla Cleome microcarpa Cleome monophylloides Cleome moricandii Cleome moritziana Cleome mossamedensis Cleome oligandra Cleome omanensis Cleome ornithopodioides Cleome oxalidea Cleome oxypetala Cleome pakistanica Cleome pallida Cleome paludosa Cleome paradoxa Cleome parviflora Cleome perrieri Cleome polyanthera Cleome polytricha Cleome potosina Cleome procumbens Cleome rosea Cleome scaposa Cleome schimperi Cleome schlechteri Cleome selloana Cleome semitetrandra Cleome serrata Cleome sessilifolia Cleome siliculifera Cleome stenopetala Cleome stenophylla Cleome stevensiana Cleome strigosa Cleome stylosa Cleome suffruticosa Cleome tenella Cleome tenuifolia Cleome tenuis Cleome titubans Cleome trachycarpa Cleome tucumanensis Cleome villosa Cleome werdermannii Cleome horrida Cleome rubelloides Cleome arenitensis Cleome limmenensis Cleome insolata Cleome bundeica Cleome lophosperma Cleome affinis Cleome ariana Cleome aspera Cleome augustinensis Cleome bojeri Cleome boliviensis Cleome brachiata Cleome brachycarpa Cleome chapalaensis Cleome chilensis Cleome chodatiana Cleome chrysantha Cleome circassica Cleome cleomoides Cleome coluteoides Cleome conrathii Cleome cordobensis Cleome cremoloba Cleome drepanocarpa Cleome droserifolia Cleome ecuadorica Cleome frutescens Cleome gallaensis Cleome gardneri Cleome gaudichaudii Cleome guianensis Cleome hadramautica Cleome hanburyana Cleome hemsleyana Cleome heratensis Cleome hirta Cleome jamesonii Cleome kalachariensis Cleome kelleriana Cleome lechleri Cleome leptorachis Cleome lilloi Cleome limoneolens Cleome linophylla Cleome lipskyi Cleome longifolia Cleome macrorhiza Cleome maculata Cleome magnifica Cleome melanosperma Cleome mexicana Cleome microaustralica Cleome niamniamensis Cleome parvipetala Cleome parvisepala Cleome parvula Cleome paxii Cleome pilosa Cleome pruriens Cleome puberula Cleome pulchella Cleome quinquenervia Cleome raddeana Cleome ramosissima Cleome regnellii Cleome rostrata Cleome rotundifolia Cleome rubella Cleome rupicola Cleome silvatica Cleome simplicifolia Cleome sinaloensis Cleome socotrana Cleome tomentella Cleome torticarpa Cleome trachysperma Cleome tunarensis Cleome turkmena Cleome uncifera Cleome uniglandulosa Cleome usambarica Cleome virens Cleome aculeata Cleome speciosa Cleome gigantea Cleome macrocarpa Cleome brachycarpa Cleome domingensis Cleome dumosa Cleome brachiata Cleome angustifolia Cleome carnosa Cleome paxii Cleome heydeana Cleome houtteana Cleome coccinea Cleome hispidula Cleome jamesii Cleome bahiensis Cleome chiriquensis Cleome puccionia Cleome erosa Cleome briquetii Cleome palustris Cleome perplexa Cleome tetrandra Cleome yunnanensis Cleome iberidella Cleome oxyphylla Cleome dumosa Cleome mathewsii Cleome refracta Cleome flava Cleome brevipetiolata Cleome crenopetala Cleome dendroidea Cleome guaranitica Cleome chelidonii Cleome afrospina Cleome albescens Cleome allamani Cleome angustifolia Cleome monophylla Cleome rutidosperma Cleome violacea Cleome viscosa Cleome gynandra Cleome spinosa Cleome bororensis Cleome stricta