Terrestrial, caulescent, erect herbs, without modified stems. Stems unbranched. Leaves opposite, nearly equal in a pair, venation pinnate, foliar nectaries absent. Flowers axillary, 1 to few in umbellate-cymose inflorescences; short pedunculate; bracteoles small; long pedicellate. Calyx lobes 5, free; corolla yellowish, tubular, inflated ventrally, limb 5-lobed; stamens included, filaments basally connate, anthers free or coherent in 2 pairs, dehiscing by longitudinal slits, thecae parallel; staminode present, small; disc of 2 or 4 opposite separate glands, or a double connate dorsal gland; ovary superior, stigma capitate or 2-lobed. Fruit a fleshy, indehiscent, black berry.