Cyperus difformis L.

Variable flatsedge (en), Souchet difforme (fr)

Species

Angiosperms > Poales > Cyperaceae > Cyperus

Characteristics

Annual with fibrous, reddish roots. Stems rather weak, tufted, triquetrous, smooth, 10-40(-65) cm by (1-)2-3 mm. Leaves flaccid, slightly canaliculate, rather abruptly acuminate, smooth or somewhat scaberulous on the margins at the top, 2-5 mm wide; lower sheaths stramineous to brown. Inflorescence simple or compound, usually rather lax, 1½-5(-7) cm across. Involucral bracts 2-3, patent, but the lowest often suberect (and then the inflorescence seemingly lateral), the 2 largest far overtopping the inflorescence, up to 25 cm. Primary rays usually 5-9, spreading, smooth, up to 3(-5) cm, secondary ones when present up to 1 cm. Spikes globose, very dense, with very numerous spikelets, 8-15 (rarely 3-4) mm across. Spikelets stellately spreading, linear or oblong-linear, compressed but slightly turgid, obtuse, 10-30-flowered, 2½-5 by 1-1¼ mm; rachilla straight, wingless, persistent; internodes c. 1/5 mm. Glumes very small, membranous, patulous, slightly keeled, I orbicular to broadly obovate, very obtuse or somewhat emarginate, muticous, with arcuate, green, faintly 3-nerved keel, purplish or sometimes pale, nerveless sides and broad whitish hyaline margins, muticous, 3/5-4/5 by ¾-1 mm, ½ imbricate. Stamens (1-)2; anthers elliptic or elliptic-oblong, c. 1/10 mm.; Style very short, 1/10-1/5 mm; stigmas 3, somewhat longer than the style. Nut triquetrous, ellipsoid to slightly obovoid, broadly stipitate, conically apiculate, about as long as the subtending glume, shining stramineous or pale brown, ½-⅔ by c. ⅓ mm.
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Herbs, annual, cespitose. Culms 1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous. Leaves 2–7, flat, (2–)7–22 cm × 2.2–4 mm. Inflorescences: heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.; rays 1–5, 2–32 mm; bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous. Spikelets 30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm; floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate. Flowers: stamens 1 or 2; anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged; styles 0.1 mm; stigmas 0.1–0.3 mm. Achenes light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose.
Annuals. Roots fibrous. Culms tufted, 2-65 cm tall, slightly thick or flaccid, compressed triquetrous, smooth, few leaved at basal part. Leaves shorter than culm; sheath brown, slightly long; leaf blade 2-6 mm wide, flat or folded. Involucral bracts 2(or 3), leaflike, longer than inflorescence. Inflorescence a simple or rarely compound anthela; rays 3-9, mostly to 3(-5) cm, unequal in length, each with many spikelets at apex densely aggregated into a capitulum. Capitulum of spikelets globose, 0.5-1.5 cm in diam. Spikelets narrowly ovoid to linear, 2-8 × 1-1.2 mm, compressed orbicular, 8-28-flowered; rachilla wingless. Glumes dark reddish purple to dark grayish brown on both surfaces but middle yellowish, less than 1 mm, slightly spreading, obscurely 3-veined, margin white hyaline, apex rounded. Stamens (1 or)2; anthers ellipsoid; connective not prominent beyond anthers. Style very short; stigmas 3, short. Nutlet yellowish, obovoid-ellipsoid, nearly as long as subtending glume, 3-sided. Fl. and fr. Jul-Oct. 2n = 18, 28, 32, 34, 36.
Annual herb, tufted, 0.05-0.80 m high. Leaf blades up to 3.5 mm wide, flat, flaccid. Culm scapose, ± 3-angled, often with a lax spiral twist, 2.5-3.5 mm in diam. Inflorescence variable, ranging from a simple, digitate, head-like cluster of spikelets to a compound anthela. Subtending bracts 3 or 4, leaf-like. Spikelets linear, 5 x 0.8 mm, 10-30-flowered. Glumes minute, boat-shaped, 0.6-0.8 mm long, keel not excurrent or excurrent into minute point. Flowering time Dec.-June. Nutlet obovate-elliptic, 0.6-0.8 x 0.3-0.4 mm, 3-angled, yellowish brown, surface finely reticulate or puncticulate.
Tufted annual 2–6 dm, bearing 2–4 bracts and several sessile or short-peduncled, globose, densely fld heads 8–12 mm thick; spikelets 4–8 mm, scarcely flattened, 10–40 fld; rachilla not winged; scales obovate-rotund, often broader than long, 0.5–0.8 mm, with reddish or brownish sides; stamen 1(–3); achenes light yellowish, trigonous, about equaling the scales. Wet soil; native of the Old World tropics, intr. in se. Va.
Annual herb, up to 0.8 m high; slender. Leaf blades developed. Flowers: inflorescence a solitary, congested anthela or with many, subumbellately arranged heads; spikelets digitate; glumes and nutlets abscising individually leaving a persistent rachilla; glumes many per spikelet, 0.5-0.7 mm long, with a winged keel, dark reddish brown; Feb.-May.
A slender to medium-sized, annual herb, up to 0.8 m high. Inflorescence a solitary, congested anthela or with many, subumbellately arranged heads. Glumes 0,5-0,7 mm long with a winged keel, dark reddish brown.
A sedge. It grows 50 cm tall.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality
Pollination anemogamy
Spread barochory
Mature width (meter) -
Mature height (meter) 0.28 - 0.73
Root system fibrous-root
Rooting depth (meter) -
Root diameter (meter) 0.5
Flower color -
Blooming months -
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

In very wet places: open grasslands, river-banks, etc., but especially a characteristic weed of wet rice-fields, often associated with C. halpan, C. iria, and C. sanguinolentus, 0-1400 m (according to;, MERRILL 2000 m); flowering throughout the year.
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It is a tropical plant. It grows in marshes and open wet areas.
Light 4-9
Soil humidity 6-10
Soil texture 1-2
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

Uses animal food gene source medicinal
Edible -
Therapeutic use Diarrhea (leaf)
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 25 - 30
Germination temperacture (C°) 21 - 23
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Cyperus difformis leaf picture by Jc Mauzac (cc-by-sa)
Cyperus difformis leaf picture by Taylor Shook (cc-by-sa)
Cyperus difformis leaf picture by Diane Tran (cc-by-sa)

Flower

Cyperus difformis flower picture by Skyler (cc-by-sa)
Cyperus difformis flower picture by Matthew Horrigan (cc-by-sa)
Cyperus difformis flower picture by kristian binar (cc-by-sa)

Fruit

Cyperus difformis fruit picture by akbar sulaiman (cc-by-sa)
Cyperus difformis fruit picture by Augustin Soulard (cc-by-sa)
Cyperus difformis fruit picture by Pascal DUPUIS (cc-by-sa)

Distribution

Cyperus difformis world distribution map, present in Afghanistan, Angola, Åland Islands, Albania, Andorra, Australia, Benin, Burkina Faso, Bangladesh, Bulgaria, Bolivia (Plurinational State of), Brazil, Botswana, Central African Republic, Chile, China, Congo, Cook Islands, Comoros, Cabo Verde, Costa Rica, Dominica, Ecuador, Egypt, Spain, Ethiopia, Fiji, France, Gabon, Georgia, Ghana, Guinea, Gambia, Guinea-Bissau, Greece, Guyana, Haiti, Indonesia, India, Iran (Islamic Republic of), Iraq, Iceland, Italy, Jamaica, Japan, Kazakhstan, Kenya, Kyrgyzstan, Liberia, Libya, Sri Lanka, Lesotho, Morocco, Madagascar, Mali, Myanmar, Mozambique, Mauritania, Mauritius, Namibia, Norfolk Island, Nigeria, Nicaragua, Nepal, Pakistan, Panama, Peru, Philippines, Palau, Papua New Guinea, Puerto Rico, Korea (Democratic People's Republic of), Portugal, Réunion, Romania, Rwanda, Sudan, Senegal, Sierra Leone, Somalia, eSwatini, Seychelles, Turks and Caicos Islands, Chad, Togo, Thailand, Tokelau, Timor-Leste, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Ukraine, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Viet Nam, Yemen, South Africa, Zambia, and Zimbabwe

Conservation status

Cyperus difformis threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:304311-1
WFO ID wfo-0000371697
COL ID 33C3B
BDTFX ID 20723
INPN ID 93918
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Cyperus lateriflorus Cyperus oryzetorum Cyperus subrotundus Cyperus protractus Cyperus goeringii Cyperus holoschoenoides Cyperus viridis Cyperus difformis var. breviglobosus Cyperus difformis var. subdecompositus Cyperus difformis f. humilis Cyperus difformis f. maximus Cyperus difformis