Cyperus L.

Papyrus sedges (en), Souchet (fr)

Genus

Angiosperms > Poales > Cyperaceae

Characteristics

Perennial or annual herbs, tufted or with creeping rhizome. Stems erect or obliquely erect, solid, triquetrous, trigonous, or sometimes subterete, usually leafy only at the base, rarely halfway up. Leaves tristichous, narrowly linear, grass-like, rarely lanceolate or elliptic, the lower ones often scale-like, covering the base of the stem and the rhizome, rarely all reduced to their sheaths. Inflorescence terminal, often anthelate (terminal spike or cluster overtopped by the lateral ones), simple to decompound, umbel-like, not rarely capitate by sup-pression of the rays, in a few spp. truly umbellate. Rays subtended by a foliaceous bract similar to the leaves, the base enclosed in a tubular, two-keeled prophyll (cladoprophyll; see below, note), the bracts usually approximate as though forming an involucre. Spikelets more or less compressed, quadrangular, or subterete, 1-many-flowered; axis (rachilla) often winged by the decurrent base of the glumes, persistent or caducous (in the latter case spikelets falling off as a whole). Glumes distichous, rarely nearly so, usually 2 basal ones empty (bracteole and prophyll), rarely 3 or 4 so. Flowers bisexual, the uppermost of the spikelet often male or barren by reduction; very rarely male and female flowers in different individuals (dioecious; not in Malesia). Hypogynous bristles or scales absent. Stamens 3,2, or 1, the median one anterior, the others transversal; connective often produced into an apical appendage. Style continuous with the ovary, not or hardly incrassate at the base, caducous, 3-fid or 2-fid, very rarely almost undivided. Nut sessile or shortly stipitate, trigonous or lenticular (planoconvex, concave-convex, or biconvex).
More
Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous. Culms solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles. Leaves usually basal; ligules absent; blades keeled abaxially, flat, V-, or inversely W-shaped in cross section. Inflorescences terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet; spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts; involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike. Spikelets: scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous). Flowers bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings; perianth absent; stamens 1–3; styles linear, 2–3-fid, base deciduous or persistent; stigmas 2–3. Achenes biconvex, flattened, or trigonous.
Herbs, annual or perennial, with rhizomes or stolons. Culms caespitose or solitary, erect, usually 3-angled or acutely triquetrous, sometimes compressed triquetrous, stout or slender, leafy at basal part or at base. Leaves basal, 3-ranked, rarely without a blade; ligule absent. Involucral bracts leaflike, usually 2-10(-20 or more), at base of inflorescence. Inflorescences terminal, a simple or compound anthela, with several to many rays and usually with raylets (secondary rays); rays and raylets unequal or rarely equal in length or inflorescence capitate. Spikelets few to many in spikes, digitate clusters, or a capitulum at apex of rays, rarely single; rachilla not articulate at base, persistent, usually with narrow wings. Glumes distichous, rarely spirally imbricate; basal 1 or 2 glumes without a flower; remaining glumes each with a bisexual flower. Perianth bristles or scales absent. Stamens (1-)3. Style base not swollen; stigmas (2 or)3, deciduous at maturity. Nutlet 3-sided, smooth, puncticulate, tuberculate, or rarely reticulately striate.
Scales distichous, the lowermost one empty and ± modified; fls perfect, each in the axil of a scale; perianth wanting; stamens 3, less often 1 or 2; style 2–3-cleft, the beakless (or nearly beakless) achene accordingly lenticular or ± trigonous; spikelets few–many in dense or loose spikes or heads, each subtended by 2 small bracts; spikes or heads commonly in a simple or compound terminal umbel that is subtended by sheathless, leafy invol bracts; each ray of the umbel surrounded at base by a tubular prophyll; herbs with solid, ± triangular stems, the lvs with closed sheath and usually an elongate, grasslike blade. 600+, cosmop. Our spp. belong to 5 subgenera, as in the following key. Spp. 14–22 and 23–28 appear to be intergradient, presumably reflecting extensive hybridization.
Annual, or perennial, often rhizomatous herbs. Stems leafy at base or leaves reduced to basal sheaths. Inflorescence umbellate or capitate, surrounded by 1-several leaf-like involucral bracts. Spikelets compressed or subterete, 1-many-flowered, usually clustered in spikes at tips of rays or in a sessile spike. Flowers hermaphrodite. Glumes distichous, usually all fertile. Hypogynous bristles 0. Stamens 2-3, or 1. Style 2-3-fid, continuous with ovary, not thickened at base. Nut trigonous or biconvex. A tropical and warm temperate genus of c. 550 spp., one endemic in N.Z.; 13 adventive.
Infl. a terminal umbel-like corymb or reduced to a single head or spikelets. Spikelets compressed or subterete, 1-∞-fld; fls hermaphrodite or the terminal 1–3 occ. male. Glumes distichous. Hypog. bristles 0. Stamens 2–3, or 1. Style 2–3-fid, continuous with ovary, not thickened at base. Nut triquetrous or plano-convex. Annual or perennial, often rhizomatous herbs. Stems lfy at base, or lvs reduced to sheaths. A tropical and warm temperate genus of c. 550 spp.; the N.Z. sp. endemic.
Annuals or perennials, the culms simple, usually triangular and leafy; inflores-cence involucrate in dense spikes or in clusters, capitate or on rays which are often compound; spikelets flat or subterete, few-or many-flowered, the rachis often winged, the scales concave, 2-ranked; flowers perfect; perianth none; stamens 1-3; style 2-to 3-cleft; achene lenticular or 3-angulate.
Life form
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Most spp. are hygrophilous; they grow in moist or wet localities at low and medium altitudes. Only a few ascend to above 1600 m. The spp. of sect. Diffusi prefer moist forests. C. stoloniferus, C. radians, C. dubius, and C. pedunculatus grow on sandy sea-shores and in sandy dunes, those with creeping stolons often acting as sandbinders. C. procerus and especially C. malaccensis prefer muddy places within the influence of salt or brackish water. C. iria, C. difformis, C. pulcherrimus, C. halpan, and C. tenuispica are characteristic of inundated rice-fields, C. rotundus is a troublesome weed in gardens and on road-sides.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 6-12

Usage

Uses. Economically the genus is unimportant. The dried stems of some spp. (e.g. C. elatus, C. malaccensis, C. procerus) are used for making mats or as strings. C. brevifolius, C. kyllingia, and C. rotundus make rather good pasturage when they are young and grow extensively. C. flabelliformis and C. papyrus are grown as ornamentals in ponds. For C. flabelliformis see p. 618.Cyperus papyrus L., indigenous in Central tropical Africa, is readily recognizable by its up to 5 m tall, pithy stems at the base covered with bladeless sheaths, the umbellate inflorescence with very numerous, subequal rays, and the lanceolate involucral bracts much shorter than the inflorescence. It was early used for paper-making and cultivated for this purpose in Egypt, Palestine, and S. Europe. See CHIOVENDA Mem. Istit. Bot. Modena 1 1931 1-120 t. 1-4 .The tubers of C. esculentus and C. bulbosus are edible, but their use is not recorded for Malesia.
Uses medicinal ornamental
Edible -
Therapeutic use Chaplet (unspecified), Necklet (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 25 - 30
Germination temperacture (C°) 21 - 23
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Cyperus unspecified picture
Cyperus unspecified picture

Distribution

Cyperus world distribution map, present in Aruba, Afghanistan, Angola, Åland Islands, Albania, Andorra, Austria, Benin, Burkina Faso, Bangladesh, Bulgaria, Bahamas, Belarus, Belize, Bermuda, Bolivia (Plurinational State of), Botswana, Central African Republic, Switzerland, Chile, China, Congo, Cook Islands, Colombia, Comoros, Cabo Verde, Costa Rica, Cuba, Cayman Islands, Cyprus, Czech Republic, Germany, Djibouti, Dominica, Denmark, Algeria, Ecuador, Egypt, Eritrea, Spain, Ethiopia, Fiji, France, Micronesia (Federated States of), Gabon, Georgia, Ghana, Guinea, Gambia, Guinea-Bissau, Greece, Guatemala, Guyana, Honduras, Haiti, Hungary, Indonesia, Ireland, Iran (Islamic Republic of), Iraq, Italy, Jamaica, Japan, Kazakhstan, Kenya, Kyrgyzstan, Kuwait, Lao People's Democratic Republic, Liberia, Libya, Lesotho, Morocco, Moldova (Republic of), Madagascar, Maldives, Mali, Myanmar, Montenegro, Mozambique, Mauritania, Mauritius, Malaysia, Namibia, Norfolk Island, Nigeria, Nicaragua, Netherlands, Nepal, Nauru, New Zealand, Oman, Pakistan, Panama, Pitcairn, Peru, Philippines, Palau, Papua New Guinea, Poland, Puerto Rico, Korea (Democratic People's Republic of), Portugal, Paraguay, Réunion, Romania, Russian Federation, Rwanda, Saudi Arabia, Sudan, Senegal, Singapore, Solomon Islands, Sierra Leone, Somalia, Suriname, Sweden, eSwatini, Seychelles, Turks and Caicos Islands, Chad, Togo, Thailand, Tokelau, Turkmenistan, Tonga, Tunisia, Tuvalu, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Ukraine, Uruguay, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Viet Nam, Vanuatu, Wallis and Futuna, Samoa, Yemen, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:330001-2
WFO ID wfo-4000010449
COL ID 8VWNX
BDTFX ID 84884
INPN ID 191454
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Hypaelyptum Volkiella Juncellus Mariscus Cyperus Torulinium

Lower taxons

Cyperus acuminatus Cyperus acuticarinatus Cyperus afromontanus Cyperus afrovaricus Cyperus albiceps Cyperus albopurpureus Cyperus albostriatus Cyperus alopecuroides Cyperus alterniflorus Cyperus altochrysocephalus Cyperus altomicroglumis Cyperus amauropus Cyperus amuricus Cyperus anderssonii Cyperus aquatilis Cyperus assimilis Cyperus astartodes Cyperus aster Cyperus aureoalatus Cyperus aureovillosus Cyperus baronii Cyperus barrosianus Cyperus benadirensis Cyperus berroi Cyperus bifolius Cyperus bipartitus Cyperus breedlovei Cyperus brumadoi Cyperus brunneofibrosus Cyperus burkartii Cyperus camphoratus Cyperus canus Cyperus castaneus Cyperus cataractarum Cyperus cearaensis Cyperus cellulosoreticulatus Cyperus centralis Cyperus cephalanthus Cyperus cephalotes Cyperus ceylanicus Cyperus chlorotropis Cyperus chordorrhizus Cyperus ciliatus Cyperus clarus Cyperus clavinux Cyperus colchicus Cyperus colymbetes Cyperus comosipes Cyperus compactus Cyperus concinnus Cyperus confertus Cyperus congensis Cyperus conglomeratus Cyperus conicus Cyperus constanzae Cyperus coriifolius Cyperus corymbosus Cyperus costaricensis Cyperus cracens Cyperus crassipes Cyperus cremeomariscus Cyperus crispulus Cyperus croceus Cyperus crypsoides Cyperus cuspidatus Cyperus cyperinus Cyperus dactylotes Cyperus x deamii o'neill Cyperus delavayi Cyperus densibulbosus Cyperus dentatus Cyperus dentoniae Cyperus dichromus Cyperus dichrostachyus Cyperus dietrichiae Cyperus digitatus Cyperus diloloensis Cyperus dipsaceus Cyperus dives Cyperus drummondii Cyperus dubius Cyperus duripes Cyperus echinatus Cyperus elegans Cyperus entrerianus Cyperus eremicus Cyperus exaltatus Cyperus fauriei Cyperus fendlerianus Cyperus ferrugineoviridis Cyperus fertilis Cyperus fibrillosus Cyperus fissus Cyperus flaccidus Cyperus flavoculmis Cyperus flexuosus Cyperus floribundus Cyperus floridanus Cyperus fugax Cyperus fulgens Cyperus fulvus Cyperus fuscescens Cyperus gardneri Cyperus giganteus Cyperus gigantobulbes Cyperus glaber Cyperus glomeratus Cyperus gossweileri Cyperus gracilis Cyperus gracillimus Cyperus granatensis Cyperus grandibulbosus Cyperus granitophilus Cyperus grayi Cyperus grayioides Cyperus guatemalensis Cyperus guianensis Cyperus gunnii Cyperus gymnocaulos Cyperus haematodes Cyperus hakonensis Cyperus harrisii Cyperus haspan Cyperus hayesii Cyperus helferi Cyperus hemisphaericus Cyperus hensii Cyperus hermaphroditus Cyperus hilairenus Cyperus hillebrandii Cyperus holoschoenus Cyperus houghtonii Cyperus humilis Cyperus hyalinus Cyperus hypochlorus Cyperus hypopitys Cyperus hystricinus Cyperus impolitus Cyperus inauratus Cyperus infucatus Cyperus inops Cyperus insidiosus Cyperus intricatus Cyperus involucratus Cyperus kappleri Cyperus karlschumannii Cyperus kasamensis Cyperus kersteinii Cyperus kipasensis Cyperus kirkii Cyperus kituiensis Cyperus kwaleensis Cyperus laeteflorens Cyperus laevigatus Cyperus laevis Cyperus lancastriensis Cyperus laxus Cyperus leptocladus Cyperus ligularis Cyperus longifolius Cyperus longi-involucratus Cyperus longistylus Cyperus luerssenii Cyperus lupulinus Cyperus luteus Cyperus macer Cyperus macropachycephalus Cyperus macrophyllus Cyperus macrorrhizus Cyperus malaccensis Cyperus maranguensis Cyperus marlothii Cyperus marojejyensis Cyperus melanospermus Cyperus melas Cyperus meyenianus Cyperus michoacanensis Cyperus micrantherus Cyperus microaureus Cyperus microbrunneus Cyperus microiria Cyperus micromelas Cyperus microstylus Cyperus microumbellatus Cyperus miliifolius Cyperus molliglumis Cyperus moutona Cyperus mundtii Cyperus mundulus Cyperus nanus Cyperus neokunthianus Cyperus niger Cyperus nigricans Cyperus nigrofuscus Cyperus nipponicus Cyperus noeanus Cyperus oakportensis Cyperus obbiadensis Cyperus oblongoincrassatus Cyperus obtusus Cyperus ochraceus Cyperus ovatus Cyperus pallidicolor Cyperus pearcei Cyperus pectinatus Cyperus pelophilus Cyperus pennatiformis Cyperus persquarrosus Cyperus pertenuis Cyperus phleoides Cyperus picardae Cyperus pinetorum Cyperus platycaulis Cyperus platyphyllus Cyperus pohlii Cyperus prieurianus Cyperus procerus Cyperus prolifer Cyperus prolixus Cyperus psammophilus Cyperus pseudothyrsiflorus Cyperus pseudovegetus Cyperus pseudovestitus Cyperus pulchellus Cyperus pulcher Cyperus pulcherrimus Cyperus pulicaris Cyperus puncticulatus Cyperus purpureoviridis Cyperus rapensis Cyperus recurvispicatus Cyperus reflexus Cyperus refractus Cyperus regiomontanus Cyperus rehmii Cyperus remotus Cyperus retrofractus Cyperus rhynchosporoides Cyperus richardii Cyperus ridleyi Cyperus rockii Cyperus rubicundus Cyperus rubriglumosus Cyperus sandwicensis Cyperus sanguineoater Cyperus scaber Cyperus scabricaulis Cyperus schaffneri Cyperus schimperianus Cyperus schomburgkianus Cyperus sciaphilus Cyperus sculptus Cyperus seemannianus Cyperus semifertilis Cyperus semiochraceus Cyperus semitrifidus Cyperus serotinus Cyperus seslerioides Cyperus sexangularis Cyperus sikkimensis Cyperus silletensis Cyperus simplex Cyperus smithianus Cyperus solidifolius Cyperus solidus Cyperus sordidus Cyperus spectabilis Cyperus sphaeroideus Cyperus sphaerolepis Cyperus spicigerus Cyperus stradbrokensis Cyperus subbadius Cyperus subcastaneus Cyperus subfuscus Cyperus subtenax Cyperus subulatus Cyperus sylvestris Cyperus szechuanensis Cyperus tanyphyllus Cyperus tenax Cyperus tenerrimus Cyperus tenuiculmis Cyperus tenuis Cyperus tenuispica Cyperus tetracarpus Cyperus tetragonus Cyperus tetraphyllus Cyperus thunbergii Cyperus thyrsiflorus Cyperus trachysanthos Cyperus trailii Cyperus trichodes Cyperus trigonellus Cyperus trinervis Cyperus trisulcus Cyperus x turbatus Cyperus turrillii Cyperus tweediei Cyperus ugogensis Cyperus undulatus Cyperus unioloides Cyperus urbanii Cyperus usitatus Cyperus ustulatus Cyperus varicus Cyperus viscidulus Cyperus vorsteri Cyperus xerophilus Cyperus zollingeri Cyperus zollingerioides Cyperus aethiops Cyperus affinis Cyperus ajax Cyperus albus Cyperus alleizettei Cyperus almensis Cyperus alulatus Cyperus amabilis Cyperus x amuricocompressus Cyperus angolensis Cyperus anisitsii Cyperus ankaizinensis Cyperus ankaratrensis Cyperus antillanus (kük.) o'neill Cyperus appendiculatus Cyperus arenarius Cyperus armstrongii Cyperus aschenbornianus Cyperus aterrimus Cyperus atkinsonii Cyperus atribulbus Cyperus atrobrunneus Cyperus atronervatus Cyperus atrorubidus Cyperus aucheri Cyperus aureobrunneus Cyperus aureostramineus Cyperus aurifer Cyperus austrochrysanthus Cyperus babakan Cyperus balfourii Cyperus baoulensis Cyperus bellus Cyperus bernieri Cyperus betafensis Cyperus betschuanus Cyperus beyrichii Cyperus blakeanus Cyperus blastophorus Cyperus blysmoides Cyperus bowmanii Cyperus breviculmis Cyperus brevifolioides Cyperus brevifolius Cyperus buchholzii Cyperus camagueyensis Cyperus cancrorum Cyperus capensis Cyperus carinatus Cyperus castaneobellus Cyperus chaetophyllus Cyperus chalaranthus Cyperus chamaecephalus Cyperus chermezonii Cyperus chersinus Cyperus chinsalensis Cyperus chionocephalus Cyperus cinereobrunneus Cyperus clandestinus Cyperus clarkei Cyperus congestus Cyperus consors Cyperus cornelii-ostenii Cyperus cruentus Cyperus cuanzensis Cyperus cunninghamii Cyperus cymulosus Cyperus deciduus Cyperus denudatus Cyperus diamantinus Cyperus dichromeniformis Cyperus diffusus Cyperus dioicus Cyperus distans Cyperus distinctus Cyperus djurensis Cyperus divulsus Cyperus drakensbergensis Cyperus dregeanus Cyperus duclouxii Cyperus elephantinus Cyperus enervis Cyperus fastigiatus Cyperus felipponei Cyperus filiculmis Cyperus filifolius Cyperus filiformis Cyperus filipes Cyperus fischerianus Cyperus fontinalis Cyperus friburgensis Cyperus fuligineus Cyperus fulvoalbescens Cyperus fuscovaginatus Cyperus glaucophyllus Cyperus graciliculmis Cyperus grammicus Cyperus grandifolius Cyperus grandis Cyperus grandisimplex Cyperus grossianus Cyperus gypsophilus Cyperus haematocephalus Cyperus hamulosus Cyperus hieronymi Cyperus holostigma Cyperus imbecillis Cyperus impubes Cyperus inaequalis Cyperus indecorus Cyperus isabellinus Cyperus ischnos Cyperus juncelliformis Cyperus kurzii Cyperus lacunosus Cyperus latifolius Cyperus latzii Cyperus lecontii Cyperus lehmannii Cyperus leiocaulon Cyperus lentiginosus Cyperus leptorhachis Cyperus linearispiculatus Cyperus lipocarphoides Cyperus longivaginans Cyperus lucidus Cyperus macranthus Cyperus maculatus Cyperus marginatus Cyperus marquisensis Cyperus matudae Cyperus medusaeus Cyperus megalanthus Cyperus megapotamicus Cyperus melanacme Cyperus meridionalis Cyperus x mesochorus Cyperus meyerianus Cyperus microbulbosus Cyperus microglumis Cyperus mogadoxensis Cyperus mudugensis Cyperus muricatus Cyperus mutisii Cyperus mwinilungensis Cyperus myrmecias Cyperus nanellus Cyperus niederleinianus Cyperus nitidus Cyperus niveoides Cyperus nuerensis Cyperus nutans Cyperus nyassensis Cyperus obtusatus Cyperus ohwii Cyperus onerosus Cyperus orgadophilus Cyperus orthostachyus Cyperus overlaetii Cyperus owanii Cyperus pachyrhizus Cyperus pagotii Cyperus palianparaiensis Cyperus pandanophyllum Cyperus pangorei Cyperus paniceus Cyperus pannonicus Cyperus paolii Cyperus pedunculosus Cyperus pendulus Cyperus penzoanus Cyperus perangustus Cyperus pernambucensis Cyperus pervillei Cyperus pilosulus Cyperus pilosus Cyperus planifolius Cyperus platystylis Cyperus plukenetii Cyperus pluribracteatus Cyperus pluricephalus Cyperus podocarpus Cyperus poecilus Cyperus poeppigii Cyperus polyanthelus Cyperus portae-tartari Cyperus praealtus Cyperus praemorsus Cyperus pratensis Cyperus pratorum Cyperus pseudobrunneus Cyperus pseudohildebrandtii Cyperus pseudokyllingioides Cyperus pseudopilosus Cyperus pumilus Cyperus radians Cyperus ramosus Cyperus retroflexus Cyperus retrorsus Cyperus rigens Cyperus rigidellus Cyperus rigidifolius Cyperus robinsonii Cyperus rohlfsii Cyperus rupicola Cyperus schinzii Cyperus schlechteri Cyperus schweinitzii Cyperus sellowianus Cyperus sensilis Cyperus setiformis Cyperus setigerus Cyperus sharonensis Cyperus soongoricus Cyperus sphacelatus Cyperus sphaerospermus Cyperus splendens Cyperus sporobolus Cyperus squarrosulus Cyperus squarrosus Cyperus steadii Cyperus subparadoxus Cyperus sulcinux Cyperus surinamensis Cyperus swartzii Cyperus tabina Cyperus tabularis Cyperus tacnensis Cyperus tanganyicanus Cyperus thomsonii Cyperus thorelii Cyperus thorncroftii Cyperus tibialis Cyperus tomaiophyllus Cyperus tonkinensis Cyperus turrialbanus Cyperus uncinulatus Cyperus unifolius Cyperus vaginatus Cyperus vestitus Cyperus wailleyi Cyperus wallichianus Cyperus xanthostachyus Cyperus zonatus Cyperus nyererei Cyperus wilburii Cyperus wissmannii Cyperus parishii Cyperus pedunculatus Cyperus natalensis Cyperus nayaritensis Cyperus pennellii o'neill Cyperus macrostachyos Cyperus oxylepis Cyperus pacificus Cyperus perennis (m.e.jones) o'neill in Cyperus laxespicatus Cyperus limosus Cyperus microbracteatus Cyperus micromariscus Cyperus micromedusaeus Cyperus micropelophilus Cyperus malawicus Cyperus mapanioides Cyperus mangorensis Cyperus manimae Cyperus somaliensis Cyperus sphaerocephalus Cyperus scleropodus Cyperus pseudosomaliensis Cyperus remotispicatus Cyperus sanguinolentus Cyperus phaeolepis Cyperus javanicus Cyperus karisimbiensis Cyperus lanceolatus Cyperus ixiocarpus Cyperus kibweanus Cyperus laetus Cyperus chrysocephalus Cyperus kaessneri Cyperus bracheilema Cyperus brunneoalbus Cyperus capillifolius Cyperus boreobellus Cyperus boreochrysocephalus Cyperus cundudoensis Cyperus aromaticus Cyperus atractocarpus Cyperus auriculatus Cyperus baobab Cyperus blepharoleptos Cyperus gayi Cyperus hilgendorfianus Cyperus incompressus Cyperus elatus Cyperus flavescens Cyperus flavidus Cyperus ekmanii Cyperus erectus Cyperus filicinus Cyperus foliaceus Cyperus cyprius Cyperus demangei Cyperus dwarkensis Cyperus imbricatus Cyperus koyaliensis Cyperus lateriticus Cyperus margaritaceus Cyperus nduru Cyperus pustulatus Cyperus reduncus Cyperus submicrolepis Cyperus spissiflorus Cyperus algeriensis Cyperus ephemerus Cyperus forskalianus Cyperus gubanii Cyperus rheophyticus Cyperus simaoensis Cyperus diwakarii Cyperus limiticola Cyperus karthikeyanii Cyperus alvesii Cyperus muniziae Cyperus davidsei Cyperus pseudopetiolatus Cyperus alaticaulis Cyperus eboracensis Cyperus unispicatus Cyperus absconditicoronatus Cyperus monospermus Cyperus hainanensis Cyperus insularis Cyperus longispicula Cyperus mauretaniensis Cyperus pseuderemicus Cyperus pulguerensis Cyperus sahelii Cyperus subtenuis Cyperus microcristatus Cyperus vandervekenii Cyperus sesquiflorus Cyperus acholiensis Cyperus unicolor Cyperus incomtus Cyperus jeminicus Cyperus nigripes Cyperus niigatensis Cyperus lacustris Cyperus tenuifolius Cyperus brunneus Cyperus stoloniferus Cyperus muliifolius Cyperus tetraformis Cyperus badius Cyperus michelianus Cyperus michelianus Cyperus cyperoides Cyperus potiguar Cyperus salzmannianus Cyperus stergiosii Cyperus inselbergensis Cyperus beentjei Cyperus abietinus Cyperus crassicuspis Cyperus hemidrummondii Cyperus hemioccidentalis Cyperus isolepis Cyperus leptocarpus Cyperus leucaspis Cyperus lipoater Cyperus lipocomosus Cyperus lipofiliformis Cyperus lipomexicanus Cyperus lipomonostachyus Cyperus lipopygmaeus Cyperus liporobinsonii Cyperus lipothermalis Cyperus purpureoluteus Cyperus raynalianus Cyperus reddyi Cyperus schomburgkii Cyperus subsquarrosus Cyperus ampullaceus Cyperus ascocapensis Cyperus ascodensus Cyperus ascofibrillosus Cyperus ascohemisphaericus Cyperus asconeglectus Cyperus ascopinguis Cyperus ascopusillus Cyperus ascospinulosus Cyperus ascotrigonus Cyperus brasiliensis Cyperus eriocauloides Cyperus erythrocephalus Cyperus majestuosus Cyperus metallorum Cyperus pseudopeteri Cyperus proteus Cyperus siamensis Cyperus vatkeanus Cyperus afro-occidentalis Cyperus auratus Cyperus carinalaevis Cyperus cataphyllatus Cyperus chrysanthoides Cyperus ngothe Cyperus ruwenzoriensis Cyperus serratangulus Cyperus africanus Cyperus acaulescens Cyperus neocooperi Cyperus okavangensis Cyperus paramoensis Cyperus conservator-davidii Cyperus hortensis Cyperus coonoorensis Cyperus stewartii Cyperus volkielloides Cyperus albescens Cyperus mahadevanii Cyperus pyramidalis Cyperus munnarensis Cyperus tuckerianus Cyperus scott-elliotii Cyperus nyikanus Cyperus kamtschaticus Cyperus beninensis Cyperus eglandulosus Cyperus kanarensis Cyperus aristulatus Cyperus kernii Cyperus mindorensis Cyperus neochinensis Cyperus nigriceps Cyperus blatteri Cyperus brunneoalatus Cyperus cardosoi Cyperus constrictus Cyperus distichus Cyperus palmatus Cyperus poikilostachys Cyperus rukwanus Cyperus scaettae Cyperus shepherdii Cyperus mbitheanus Cyperus hooperae Cyperus afrorobustus Cyperus breviglumis Cyperus camerunensis Cyperus felicis Cyperus kilianii Cyperus pseudobulbosus Cyperus robinsonianus Cyperus tisserantioides Cyperus xantholepis Cyperus austroafricanus Cyperus decurvatus Cyperus uitenhagensis Cyperus steudneri Cyperus atriceps Cyperus betchei Cyperus bulbipes Cyperus caesius Cyperus cartilagineus Cyperus chorisanthos Cyperus ciliatopilosus Cyperus dewildeorum Cyperus durus Cyperus erinaceus Cyperus ivohibensis Cyperus kabarensis Cyperus mollipes Cyperus oxycarpus Cyperus pachystylus Cyperus panamensis Cyperus pauper Cyperus rhizomatosus Cyperus sieberianus Cyperus stenophyllus Cyperus sublaevicarinatus Cyperus svensonii Cyperus trialatus Cyperus stolonifer Cyperus afroalpinus Cyperus arsenei o'neill Cyperus compressiformis Cyperus controversus Cyperus coronarius Cyperus costatus Cyperus disjunctus Cyperus dunensis Cyperus erythrorrhizos Cyperus eximius Cyperus familiaris Cyperus hesperius Cyperus laxiflorus Cyperus lundellii o'neill Cyperus mirus Cyperus multifolius Cyperus peteri Cyperus pseudodiaphanus Cyperus substramineus Cyperus pubens Cyperus renschii Cyperus rhizomafragilis Cyperus somalidunensis Cyperus stramineoferrugineus Cyperus subaequalis Cyperus subpapuanus Cyperus subtrigonus Cyperus tanzaniae Cyperus victoriensis Cyperus zonatissimus Cyperus dipsacoides Cyperus echinus Cyperus perspicuus Cyperus altsonii Cyperus redolens Cyperus longistolon Cyperus subtilis Cyperus plantaginifolius Cyperus perrieri Cyperus soyauxii Cyperus subcaracasanus Cyperus ventricosus Cyperus sumbawangensis Cyperus sexflorus Cyperus diaphanus Cyperus matagoroensis Cyperus tatandaensis Cyperus volodia Cyperus boreohemisphaericus Cyperus brunnescens Cyperus chermezonianus Cyperus chevalieri Cyperus chlorocephalus Cyperus chrysanthus Cyperus columbiensis Cyperus eglobosus Cyperus meistostylus Cyperus membranaceus Cyperus microbolbos Cyperus microcephalus Cyperus multinervatus Cyperus multispicatus Cyperus nemoralis Cyperus nudiceps (c.b.clarke ex standl.) o'neill Cyperus welwitschii Cyperus plateilema Cyperus derreilema Cyperus malabaricus Cyperus mitis Cyperus kyllingiella Cyperus hystricoides Cyperus afropumilus Cyperus songeensis Cyperus simpsonii Cyperus monoflorus Cyperus albopilosus Cyperus albosanguineus Cyperus angustatus Cyperus holstii Cyperus neobarteri Cyperus penicillatus Cyperus pycnostachyus Cyperus rupestris Cyperus sartorii Cyperus secubans Cyperus tempeae Cyperus tenuispiculatus Cyperus textilis Cyperus whitmeei Cyperus calderoniae Cyperus celans Cyperus commixtus Cyperus correllii Cyperus cristulatus Cyperus curvistylis Cyperus dactyliformis Cyperus debilissimus Cyperus decompositus Cyperus dilatatus Cyperus endlichii Cyperus expansus Cyperus feani Cyperus fucosus Cyperus heterocladus Cyperus hirtellus Cyperus kyllingiformis Cyperus lhotskyanus Cyperus locuples Cyperus majungensis Cyperus neoguinensis Cyperus neotropicalis Cyperus neourbanii Cyperus nervosostriatus Cyperus ornatus Cyperus permacer Cyperus pseudodistans Cyperus mortonii Cyperus nervulosus Cyperus volckmannii Cyperus leucocephalus Cyperus luteostramineus Cyperus maderaspatanus Cyperus ossicaulis Cyperus pachycephalus Cyperus scariosus Cyperus schweinfurthii Cyperus plurinervosus Cyperus rufostriatus Cyperus afrodunensis Cyperus afroechinatus Cyperus albogracilis Cyperus bigibbosa Cyperus phillipsiae Cyperus alatus Cyperus gilesii Cyperus intactus Cyperus bulbosus Cyperus meeboldii Cyperus yadavii Cyperus multispiceus Cyperus sharpei Cyperus virens Cyperus aggregatus Cyperus kilimandscharicus Cyperus odoratus Cyperus luzulae Cyperus fuscus Cyperus difformis Cyperus compressus Cyperus iria Cyperus strigosus Cyperus papyrus Cyperus niveus Cyperus articulatus Cyperus alternifolius Cyperus longus Cyperus rotundus Cyperus eragrostis Cyperus esculentus Cyperus capitatus