Cyperus iria L.

Ricefield flatsedge (en)

Species

Angiosperms > Poales > Cyperaceae > Cyperus

Characteristics

Annual, or perennial in favourable circumstances; roots yellowish red. Stems tufted, slender to rather stout, triquetrous, smooth, (5-)15-50(-80) cm by (1-)2-3(-5) mm. Leaves basal, shorter than to somewhat longer than the stems, flat or channelled, rather flaccid, gradually acuminate, scabrous on the margins in the upper parts, 3-6(-8) mm wide. Inflorescence greatly varying in size, simple or compound, usually loose, up to 20 cm long. Involucral bracts 3-5(-7), obliquely erect to patent, the larger 1-3 overtopping the inflorescence, up to 40 cm long. Primary rays 3-5(-8), very unequal, obliquely patent, slender, smooth, up to 10(-18) cm, the larger ones usually umbellately or paniculately branched, the secondary rays very short. Spikes narrow, oblong-ovoid, often elongate, rather dense or loose, with 5-25 spikelets; rachis flexuous, glabrous. Spikelets spicately arranged, oblong-linear, strongly compressed, obtuse, (in the Malesian specimens) erecto-patent, 6-20(-24)-flowered, 3-10(-13) by 1½-2 mm; rachilla straight, wingless, persistent; internodes c. ½ mm. Glumes membranous, spreading-ascendent, orbicular or broadly ovate, not rarely broader than long, sharply keeled, rounded to emarginate at apex, very shortly mucronulate with green, arched, 3-5-nerved keel, cellular-reticulate, nerveless, golden to fulvous sides, and broadly whitish hyaline margins towards the top, ⅓-½ imbricate, 1¼-1½ by 1-1½ mm. Stamens 2-3; anthers very small, oblong, c. ¼ mm, with short, smooth, reddish appendage of the connective. Style hardly any; stigmas 3, ¼-½ mm long. Nut about as long as the subtending glume, triquetrous with slightly concave sides, obovoid-ellipsoid, broadly stipitate, hardly apiculate, shining dark brown to black, 1-1½ by ½-¾ mm.
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Annuals. Roots fibrous. Culms tufted, 8-80 cm tall, slender to slightly stout, compressed triquetrous, base few leaved. Leaves much shorter than culm; sheath reddish brown to brownish purple; leaf blade 2-5 mm wide, slightly folded or flat. Involucral bracts 3-5, leaflike, lowermost 2 or 3 longer than inflorescence. Inflorescence a compound or rarely simple anthela; rays 4-9, mostly to 12 cm, each with 5-10 or more spikes. Spikes ovoid to oblong-ovoid, 1-4 cm, with 5-20 or more spikelets. Spikelets laxly arranged, ascending to slightly spreading, narrowly ovoid, very narrowly ovoid, or oblong, 4-10 × ca. 2 mm, compressed, 6-22-flowered; rachilla almost wingless, with successive glumes 0.7-0.9 mm apart. Glumes yellow to straw-colored, lax, broadly obovate, ca. 1.5 mm, membranous, 3-5-veined, keel middle abaxially green, margin apically white hyaline, apex retuse and mucronulate. Stamens 2(or 3); anthers ellipsoid; connective not prominent beyond anthers. Style very short; stigmas ± short. Nutlet dark brown, obovoid to subellipsoid, 1.2-1.4 mm, nearly as long as subtending glume, 3-sided, densely prominently puncticulate. Fl. and fr. Jun-Oct. 2n = 108, 112, 116, 128.
Herbs, annual, cespitose. Culms trigonous, (5–)20–60 cm × (0.2–) 0.6–2.4 mm, glabrous. Leaves (1–)3–4, V-shaped, 15–40 cm × 3–6 mm. Inflorescences: spikes 2–5(–8), oblong to ovoid, 2–10 cm × 3–12 mm; rays 4–8, to 11 cm; bracts (4–)5(–7), ascending at 45°, V-shaped, (1.5–)8–30(–50) cm × 1–6 mm; rachilla persistent, wingless (successive floral scales 0.7–0.9 mm apart). Spikelets 10–30, compressed, linear to ellipsoid, 4–20 × 1.5–2.1 mm; floral scales deciduous, (2–)6–26, laterally brown to golden brown, edges clear, medially green, 5-ribbed, outermost ribs more noticeable than others, obovate to nearly orbiculate, 1.3–1.8 × 1.2–1.8 mm, margins emarginate, apex with mucro 0.05–0.12 mm. Flowers: stamens 2(–3); anthers (0.2–)0.3–0.4 mm, connective tips reddish, less than 0.1 mm; styles less than 0.1 mm; stigmas 0.2–0.4 mm. Achenes brown, slightly stipitate, obovoid, 1.2–1.4 × 0.5–0.7 mm, apex apiculate, surfaces puncticulate.
Tufted annual to 6 dm, with rather short, basally disposed lvs; rays usually well developed and commonly branched at the top; spikes loose and open, 1–2 cm, bearing several rather distant, ascending spikelets; spikelets 10–20-fld, to 1 cm; rachilla persistent; scales broadly obovate, rounded-emarginate, scarcely mucronate, 1.2–1.7 mm, with green midrib and golden-brown sides; rachilla wingless; stamens 2 or 3; achenes trigonous, 1–1.5 mm, blackish. Swamps and muddy ground; native of Eurasia, intr. in se. U.S. as far n. as s. Ill., Ky. and se. Va.
Annual herb, up to 0.6 m high. Inflorescence an open anthela, 15-200 x 10-200 mm, with groups of spikes sessile or on long peduncles. Glumes with an uncoloured margin and a greenish, slightly excurrent midrib, golden brown.
Obtuse, golden, very compressed spikelets in oblong, erect spikes
Annual up to 76 cm. high
Life form annual
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.35 - 0.78
Root system fibrous-root rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c4

Environment

Open, wet locations; it is a characteristic weed in lowland rice fields, but may also occur as a weed in vegetables, maize, sugar cane, groundnut and soya bean; also occurring on roadsides and river banks; at elevations up to 700 metres.
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In open wet places, on road-sides, river-banks, and especially a characteristic weed in wet rice-fields; at low altitudes up to 700 m, rarely to 1200 m.
Edge of open water.
Light 4-9
Soil humidity 7-9
Soil texture 1-2
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

Uses animal food material medicinal
Edible stems
Therapeutic use Appetite stimulants (leaf), Antirheumatic agents (root), Astringent (unspecified), Stimulant (unspecified), Stomachic (unspecified), Tonic (unspecified), Appetite stimulants (unspecified), Astringents (unspecified), General tonic for rejuvenation (unspecified), Stomach diseases (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) 25 - 30
Germination temperacture (C°) 21 - 23
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Cyperus iria leaf picture by Ken Borgfeldt (cc-by-sa)
Cyperus iria leaf picture by João Ferreira Silva Junior (cc-by-sa)
Cyperus iria leaf picture by João Ferreira Silva Junior (cc-by-sa)

Flower

Cyperus iria flower picture by Augustin Soulard (cc-by-sa)
Cyperus iria flower picture by Mumdi Pertin (cc-by-sa)
Cyperus iria flower picture by Salahaudin Maili (cc-by-sa)

Distribution

Cyperus iria world distribution map, present in Afghanistan, Åland Islands, Andorra, Australia, Benin, Bangladesh, Belize, Bolivia (Plurinational State of), Brazil, Botswana, China, Costa Rica, Cuba, Ecuador, Eritrea, Ethiopia, Fiji, Gabon, Georgia, Ghana, Gambia, Honduras, Indonesia, Iran (Islamic Republic of), Iraq, Japan, Kenya, Lao People's Democratic Republic, Moldova (Republic of), Madagascar, Mali, Myanmar, Mozambique, Mauritania, Mauritius, Malaysia, Namibia, Nigeria, Nicaragua, Nepal, Nauru, Oman, Pakistan, Panama, Peru, Philippines, Puerto Rico, Korea (Democratic People's Republic of), Réunion, Sudan, Senegal, Somalia, eSwatini, Chad, Thailand, Turkmenistan, Taiwan, Province of China, Tanzania, United Republic of, Uganda, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Viet Nam, South Africa, Zambia, and Zimbabwe

Conservation status

Cyperus iria threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:304844-1
WFO ID wfo-0000375442
COL ID 33CHV
BDTFX ID 77708
INPN ID 161524
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Cyperus iria f. flavescens Cyperus iria f. multiflora Chlorocyperus iria Cyperus chrysomelinus Cyperus diaphaniria Cyperus nangtciangensis Cyperus paniciformis Cyperus panicoides Cyperus resinosus Cyperus santonici Cyperus microlepis Cyperus iria var. paniciformis Cyperus iria var. flavescens Cyperus iria var. santonici Cyperus iria f. chrysomelinus Cyperus iria f. diaphaniria Cyperus iria var. typicus Cyperus iria