Dacryodes costata (A.W.Benn.) H.J.Lam

Tree 20-30(-40) m by 20-100 cm, rarely with small buttresses. Branchlets c. 0.5 cm thick, minutely pubescent; pith without vascular strands. Leaves (1-)2(-4)-jugate, glabrous to densely pubescent except the upper surface of the leaflets; nerves up to 17 or 18 pairs. Petioles 1-5(-6½) cm, much flattened at base, pith with few vascular strands. Leaflets broadly elliptic to oblong, 3-16 by 1.5-7.5 cm, rigid, chartaceous to subcoriaceous, underneath more or less woolly pubescent on midrib and nerves, glabrescent; base hardly oblique, broadly cuneate to rounded; apex abruptly, shortly, and bluntly acuminate; nervation prominent beneath, nerves 6-13 pairs (angle c. 60°, at the base up to 90°), strongly curved, not arching except some apical ones. Panicles terminal and often in the uppermost leaf-axils, usually branched from the base, densely woolly pubescent, flowers in clusters; ♂ ones 6.5-30 cm, much branched, branches spreading, up to 12 cm; female ones 4-21 cm, narrowly paniculate to subracemose, branches up to 4 cm. Inflorescences apparently sometimes exclusively axillary (SAN 75957). Flowers small, slightly pubescent. Calyx 0.5-1 mm high. Petals glabrous. Filaments free from the disk. Disk annular to 6-lobed, thick. Pistil in ♂ flowers very reduced. Infructescences densely woolly pubescent. Fruits somtimes ellipsoid, hardly or not oblique, 12-19 by 8-12 mm, rounded at the base, more or less obtusely acute at the apex; wall of the pyrene rather thick and hard. Cotyledons folded, 3-lobed.

Lowland primary forests, usually at elevations below 100 metres, but occasionally ascending to 500 metres. Usually found on hillsides and ridges with sandy soils.

Uses. In Borneo the timber is used for light constructions, the bole for building proahs.

Can be grown by seedlings.