Shrubs to medium-sized trees, monoecious. Indumentum of simple, solitary hairs and small, round (glandular?) scales. Twigs terete, more or less grooved. Leaves spirally arranged, without stipules, paripinnate, 2-12-jugate. Leaflets alternate to opposite, variably hairy, sparsely to sometimes densely scaly; margin subentire to sparsely, sometimes coarsely (serrate-)crenate; domatia absent or pocket-like. Inflorescences axillary, often pseudoterminal, simple and raceme-like or ± strongly branched; cymules 1-few-flowered; bracts and bracteoles densely hairy, mostly caducous. Flowers unisexual, actino-morphic. Sepals 5 (or 6), imbricate, slightly connate at the base, suborbicular to subel-liptic, about equal or the inner-and/or outermost sepal distinctly smaller, persistent under the fruit. Petals none. Disc entire, swollen, tomentose. Stamens (4-)5(-6), exserted in male flowers; filaments usually long patently hairy in the basal half; anthers (basally to) halfway dorsally attached. Ovary (sub)sessile), 2-(or 3-)celled, ± hairy, sparsely to densely scaly; style apical, shorter to little longer than the ovary, with 2 (or 3) stigmatic lines. Ovules 1 per locule, axillary. Infructescences with thickened axes. Fruits capsular, (sub)sessile, loculicidal, wall granular, inside ± fleshy with a thicker fleshy layer in a narrow to broad strip along the dissepiment, covering the inside of the valves for 30-100%. Seeds ovoid to ellipsoid, 1 or 2 (or 3) per fruit; testa pergamentaceous or a ± hard shell; sarcotesta on the ventral side cupular and covering 25-100% of the seed.
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Small trees, monoecious. Branchlets glabrous except at apex; indumentum of simple hairs. Leaves paripinnate; leaflets entire, crenate or serrulate, thinly coriaceous or ±membranous, densely reticulate-veined, usually lepidote, petiolulate or subsessile. Inflorescence axillary, spike-like, rarely branched; cymules clustered, few-flowered. Flowers regular, pedicellate. Calyx 4-or 5-partite, 2-seriate. Petals absent. Disc regular, hairy. Stamens 5; filaments filiform, hairy. Ovary 2-or 3-sulcate, slightly compressed, 2-or 3-locular; ovule 1 per locule, 1 or 2 abortive; style short. Fruit obovoid or subglobose, usually small, 1-seeded, loculicidally dehiscent; endocarp bony. Seed ellipsoidal with cupular, bilobed ventral aril at base.
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