Herbs perennial, rhizomatous. Trichomes a mixture of simple and short-stalked, submalpighiaceous. Stems decumbent or ascending. Basal leaves rosulate, simple, entire or distally 3-5-lobed. Cauline leaves sessile and attenuate at base, entire, apically dentate. Racemes several flowered, bracteate. Fruiting pedicels ascending or divaricate at base, strongly recurved and often forming a loop. Sepals oblong, spreading, glabrous or pubescent, base of lateral pair not saccate. Petals white, sometimes with pink veins, much longer than sepals; blade broadly obcordate to orbicular, apex emarginate or rarely subrounded; claw short, well differentiated from blade. Stamens 6, erect, subequal; filaments not dilated at base; anthers ovate to suboblong, obtuse at apex. Median glands absent; lateral glands 1 on each side of lateral stamen. Ovules 4 per ovary. Fruit dehiscent silicles, oblong or ovoid, terete or slightly angustiseptate, sessile; valves membranous, sparsely to densely hirsute with subsetose trichomes, rarely glabrous, rounded, smooth, wingless or with small appendages; replum strongly flattened and to 1 mm wide; septum complete; style conical; stigma capitate, entire. Seeds wingless, oblong, flattened; seed coat minutely reticulate, not mucilaginous when wetted; cotyledons accumbent.