Echinochloa crus-galli (L.) P.Beauv.

Cockspur (en), Pied-de-coq (fr), Échinochloé Pied-de-coq (fr), Échinochloa pied-de-coq (fr), Panic pied-de-coq (fr)

Species

Angiosperms > Poales > Poaceae > Echinochloa

Characteristics

Strong, tufted, bright green, rarely glaucous, summer annuals, with stout culms, usually ± prostrate at first, particularly in open situations, becoming erect at flowering. Leaf-sheath light green or purplish, chartaceous, keeled, glabrous, or occasionally with soft tubercle-based hairs just inside margin and scattered elsewhere. Ligule 0; ligular area glabrous, rarely faintly pubescent. Leaf-blade 15-50 cm × 5-15 mm, soft, flat, linear, glabrous, or sometimes with scattered soft, tubercle-based hairs, midrib distinct; margins and sometimes midrib scabrid, tapering to acute tip. Culm (20)-30-130 cm, slender to stout, often branching and compressed below, internodes glabrous. Panicle (6)-10-20 cm, erect, ± open, very variable in shape and in presence or absence of awned florets, with few to many spike-like racemes borne singly, or in pairs or clusters at nodes; rachis 3-angled, scabrid. Racemes (1)-2.5-6 cm, erect or spreading, distant or approximate, but not extremely close-set, bearing spikelets crowded in clusters of 2 or 3; rachis 3-angled, scabrid, hairs few to many, bristle-like tubercle-based hairs clustered at base; pedicels scabrid, 0.5-1 mm. Spikelets 3-5 mm, greenish or often purplish, ovate-elliptic, acute to acuminate. Glumes quite unequal, submembranous, acute to cuspidate, scabrid; lower 1.2-1.7 mm, 3-5-nerved, broadly ovate, enwrapping base of spikelet, upper 3-4 mm, 5-nerved, very convex, with bristle-like, tubercle-based hairs on nerves, very rarely slightly awned. Lower floret: lemma similar to upper glume, 5-7-nerved, flat or dorsally depressed with central nerves often obscure, nerves, especially lateral nerves, with tubercle-based spinules, apex acute, cuspidate or distinctly awned (2-25 mm) in some florets; palea < lemma, hyaline, broad, keels finely ciliate. Upper floret: lemma 2.7-3.5 mm, ovate-elliptic, very convex, whitish or yellowish, glabrous, shining, thinly crustaceous, cuspidate, apex greenish, scabrid, submembranous, deltoid; palea ≈ lemma, apex subobtuse, soft; anthers 0.5-0.7 mm, brownish yellow or greyish; caryopsis 1.2-2 × 1.4-1.8 mm, orbicular, greenish or whitish, not turgid.
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Stout, often branched from the decumbent base; sheaths glabrous; blades 5–30 mm wide; infl 1–2.5 dm, erect, with usually 15–25 appressed or spreading branches 2–4 cm (to 6 cm in var. frumentacea), the longer ones rebranched; main axis and branches of the infl beset with stout, often papillate-based setae typically as long as or longer than the spikelets; spikelets 2.8–4 mm; second glume and sterile lemma variously scabrous, hispid, or hirsute to subglabrous, but the hairs usually not papillate-based, the glume awnless or nearly so; fertile lemma 2.5–3.5 mm, ca twice as long as wide, the short, soft beak withering, somewhat inflexed, sharply differentiated from the obtuse body, marked by a line of minute bristles at base; 2n=54. Native of Eurasia, widespread in the U.S., Can., and Mex. Two vars. with us:
Tufted annual, 0.25-1.00 m high; stoloniferous; hygrophyte. Leaf blade 70-350 X 1-20 mm; ligule absent. Inflorescence elongated, 60-220 mm long, of many racemes scattered up central axis, usually with short secondary branchlets at base; raceme 20-100 mm long, spikelets untidily 2-several-rowed. Spikelets 3-4(-7) x 1-2 mm, ovate-elliptic, dorsiventrally compressed, hispid; glumes unequal; upper glume as long as spikelet; glumes and lower lemma minutely hairy, nerves with long, rigid hairs. Florets 2; lower floret sterile, lower lemma usually awnless or sometimes with awn 5-35 mm long; upper floret bisexual, lemma 2-3 mm long, hard, entire, margins inrolled and clasping only edges of palea; anther 0.7-1.1 mm long. Flowering time Jan.-Apr.
Annual. Culms coarse, erect or geniculately ascending, 20–150 cm tall. Leaf blades linear, 5–40 × 0.2–1.2 cm, usually glabrous, smooth except for scabrous margins, apex acute. Inflorescence erect, lanceolate to ovate or pyramidal, 6–22 cm; racemes 2–10 cm, usually ascending, simple or the longest with inconspicuous branchlets near the base, rachis usually with tubercle-based setae, spikelets loosely to densely crowded. Spikelets green or purplish, ovate, 2.5–4 mm, spinulose along veins; lower glume ca. 1/3 as long as spikelet, acute; lower lemma sterile, herbaceous, acuminate or extended into an awn to 3 cm; upper lemma pale brownish at maturity, elliptic, 2–3 mm. Fl. and fr. summer and autumn. 2n = 36, 48, 54, 72.
Tufted annual 250-1 000 mm high; hygrophyte; stoloniferous. Leaf blade 70-350 x 4-20 mm, ligule absent. Inflorescence 60-220 mm long, elongated; raceme 20-100 mm long, untidily 2 to several-rowed, usually with short secondary branchlets at the base, not or inconspicuously compound. Spikelet 3-4(-7) x 1 to 2 mm, ovate-elliptic; glumes and lower lemma minutely hairy, nerves with long, rigid hairs; lower floret sterile; lower lemma usually awnless, sometimes with a awn 5-35 mm long; upper lemma 2 to 3 mm long (including short herbaceous apex); anther 0.7-1.1 mm long.
An annual grass It grows 1-1.5 m tall. It can be erect or lean over. The leaves are long and narrow and flat. They are about 35 cm long and 1 cm wide. The leaves do not have hairs except for a few hairs at the base. The flower arrangement is open and up to 25 cm long by 8 cm wide. The flower arrangement is pinkish. There are 3 or 4 irregular rows. The lower flower stalks are 10 cm long and they get shorter higher up. The spikelets are 2-4.5 mm long. It does not have a ligule or row of hairs where the leaf sheath joins the stem. One plant can produce 7000 seed.
Coarse annual; culms 25–100 cm. high, erect or spreading.. Leaf-blades 7–35 cm. long, 4–20 mm. wide; ligule absent; sheaths glabrous, rarely appressed hairy.. Inflorescence linear to ovate, 6–22 cm. long, the racemes untidily 2–several-rowed, the longest 2–10 cm. long usually with short secondary branchlets at the base.. Spikelets ovate-elliptic, mostly 3–4 mm. long, hispid; lower floret barren, acuminate or with an awn up to 5 cm. long; upper lemma 2–3 mm. long, including the short herbaceous tip.
Annual, hydrophyte, stoloniferous and tufted, up to 1 m high. Leaf blades 70-350 mm long, 4-20 mm wide; ligule absent. Spikelets 3 or 4(-7) mm long, 1-2 mm wide. Inflorescence 60-220 mm long; racemes untidily 2-to several-rowed, 20-100 mm long and usually with secondary branchlets at base; lower floret sterile; lower lemma usually awnless, rarely with an awn 5-10 mm long.
Annual; up to 1 m high; hydrophyte; stoloniferous and tufted. Leaf blades 70-350 x 4-20 mm; ligule absent. Flowers: inflorescence spike-like; 60-220 mm long; racemes untidily 2-to several-rowed; 20-100 mm long; base usually with secondary branchlets; spikelets 3-4(-7) x 1-2 mm; lower floret sterile; lower lemma usually not awned; rarely with awn 5-35 mm long.
Inflorescence 6–22 cm. long, linear to ovate; racemes untidily 2-several-rowed, the longest 2–10 cm. long usually with short secondary branchlets at the base.
Culms 25–100 cm. high, erect or ascending; ligule absent; leaf laminae 62–20 mm. wide.
Superior lemma 2–3 mm. long, including the short herbaceous apex.
Inferior lemma acuminate or with an awn up to 5 cm. long.
Spikelets mostly 3–4 mm. long, ovate-elliptic, hispid.
Coarse annual.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention -
Sexuality hermaphrodite
Pollination anemogamy
Spread epizoochory
Mature width (meter) 0.15
Mature height (meter) 0.3 - 1.0
Root system fibrous-root
Rooting depth (meter) 0.5
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c4

Environment

A tropical plant. It grows in wet places and paddy fields. It grows in wetlands. It also grows in temperate places. Seed germinate most easily at 20°C. It grows in water-logged soils which are medium to heavy. It can grow in arid places. In Argentina it grows from sea level to 800 m above sea level. Tasmanian Herbarium. In Yunnan.
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Ditches roadsides and waste places in S. Europe. Wet places and rich soils in India.
Ditches roadsides and waste places in S. Europe. Wet places and rich soils in India.
Light 7-9
Soil humidity 4-8
Soil texture 1-6
Soil acidity 3-9
Soil nutriment -
Hardiness (USDA) 6-8

Usage

The young shoots are eaten as a vegetable. The seeds can be boiled and eaten. They can also be popped like corn. They can be ground into flour and used to make bread, cakes and porridge. They are used for macaroni and dumplings. They can be steamed and made in porridge. Roasted seeds are used as a coffee substitute.
Uses animal food brewing coffee substitute environmental use fodder food gene source medicinal
Edible leaves seeds shoots stems
Therapeutic use Estrogen receptor modulators (aerial part), Anti-bacterial agents (seed), Antifungal agents (seed), Carbuncle (unspecified), Hemorrhage (unspecified), Spleen (unspecified), Sore (unspecified), Splenitis (unspecified), Estrogen receptor modulators (unspecified), Hemostatics (unspecified), Mouth diseases (unspecified), Neoplasms (unspecified), Splenic diseases (unspecified), Wounds and injuries (unspecified), Hemorrhage (whole plant), Splenic diseases (whole plant)
Human toxicity -
Animal toxicity -

Cultivation

Plants are grown from seed.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) 17 - 30
Size -
Vigor -
Productivity -

Images

Habit

Echinochloa crus-galli habit picture by Alain Lagrave (cc-by-sa)
Echinochloa crus-galli habit picture by Prieta Javier (cc-by-sa)
Echinochloa crus-galli habit picture by micka wagner (cc-by-sa)

Leaf

Echinochloa crus-galli leaf picture by Victor Luyt (cc-by-sa)
Echinochloa crus-galli leaf picture by Luc Lambert (cc-by-sa)
Echinochloa crus-galli leaf picture by Markus (cc-by-sa)

Flower

Echinochloa crus-galli flower picture by Victor Luyt (cc-by-sa)
Echinochloa crus-galli flower picture by Victor Luyt (cc-by-sa)
Echinochloa crus-galli flower picture by Markus (cc-by-sa)

Fruit

Echinochloa crus-galli fruit picture by gerhard schurr (cc-by-sa)
Echinochloa crus-galli fruit picture by Ivy Le Maguer (cc-by-sa)
Echinochloa crus-galli fruit picture by Hilairet Anthéa (cc-by-sa)

Distribution

Echinochloa crus-galli world distribution map, present in Afghanistan, Åland Islands, Albania, Andorra, Austria, Bangladesh, Bulgaria, Bahamas, Belarus, Bermuda, Brazil, Botswana, Canada, Switzerland, Chile, China, Cook Islands, Colombia, Costa Rica, Cuba, Cyprus, Czech Republic, Germany, Dominica, Denmark, Algeria, Egypt, Spain, Fiji, France, Micronesia (Federated States of), Georgia, Greece, Haiti, Hungary, Indonesia, Iran (Islamic Republic of), Iraq, Italy, Jamaica, Japan, Kazakhstan, Kyrgyzstan, Lao People's Democratic Republic, Libya, Lesotho, Morocco, Madagascar, Myanmar, Montenegro, Mauritius, Malaysia, Namibia, Norfolk Island, Nicaragua, Netherlands, Nepal, New Zealand, Oman, Pakistan, Peru, Philippines, Palau, Poland, Puerto Rico, Korea (Democratic People's Republic of), Portugal, Paraguay, Réunion, Romania, Russian Federation, Saudi Arabia, Solomon Islands, eSwatini, Turks and Caicos Islands, Togo, Thailand, Turkmenistan, Tunisia, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Ukraine, Uruguay, United States of America, Uzbekistan, Venezuela (Bolivarian Republic of), Viet Nam, Yemen, and South Africa

Conservation status

Echinochloa crus-galli threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:399660-1
WFO ID wfo-0000865236
COL ID 38GZJ
BDTFX ID 23376
INPN ID 95671
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Echinochloa crus-galli f. zelayensis Echinochloa caudata Echinochloa crus-corvi Echinochloa hispida Echinochloa macrocorvi Echinochloa madagascariensis Echinochloa micans Echinochloa occidentalis Echinochloa persistentia Echinochloa spiralis Ischaemum glabrescens Milium crus-galli Oplismenus crus-galli Oplismenus echinatus Orthopogon echinatus Panicum crista-galli Panicum crus-corvi Panicum crus-galli Panicum limosum Panicum oryzetorum Panicum scindens Echinochloa tzvelevii Echinochloa formosensis Echinochloa paracorvi Oplismenus limosus Oplismenus zelayensis Panicum alectorocnemum Panicum coarctatum Panicum echinatum Panicum goiranii Echinochloa crus-galli f. atra Echinochloa crus-galli f. breviseta Echinochloa crus-galli f. echinata Echinochloa crus-galli f. exigua Echinochloa crus-galli f. longiseta Echinochloa crus-galli f. mitis Echinochloa crus-galli f. mixta Echinochloa crus-galli f. mutica Echinochloa crus-galli f. purpurea Echinochloa disticha Echinochloa dubia Echinochloa echinata Echinochloa glabrescens Oplismenus dubius Panicum corvi Panicum corvipes Panicum crus-galli f. mixtum Panicum cruspici Panicum dubium Panicum hispidum Panicum numidianum Panicum segetale Panicum zeylayense Pennisetum crus-galli Panicum crus-galli f. atra Panicum crus-galli f. rohlenae Echinochloa crus-galli f. breviseta Oplismenus crus-corvi Panicum grossum Panicum alectromerum Orthopogon crus-galli Echinochloa commutata Echinochloa crus-galli subsp. caudata Echinochloa crus-galli subsp. spiralis Echinochloa crus-galli subsp. zelayensis Echinochloa crus-galli var. aristata Echinochloa crus-galli var. austrojaponensis Echinochloa crus-galli var. breviseta Echinochloa crus-galli var. caudata Echinochloa crus-galli var. echinata Echinochloa crus-galli var. formosensis Echinochloa crus-galli var. kasaharae Echinochloa crus-galli var. longiseta Echinochloa crus-galli var. mitis Echinochloa crus-galli var. mutica Echinochloa crus-galli var. praticola Echinochloa crus-galli var. submutica Echinochloa crus-galli var. zelayensis Echinochloa crus-pavonis var. austrojaponensis Echinochloa crus-pavonis var. breviseta Echinochloa crus-pavonis var. praticola Echinochloa macrocarpa var. aristata Echinochloa muricata var. occidentalis Echinochloa pungens var. occidentalis Oplismenus crusgalli var. muticus Panicum crus-galli var. angustifolium Panicum crus-galli var. aristatum Panicum crus-galli var. brevisetum Panicum crus-galli var. echinatum Panicum crus-galli var. longisetum Panicum crus-galli var. longisetum Panicum crus-galli var. mite Panicum crusgalli var. muticum Panicum crus-galli var. prostratum Panicum crus-galli var. pumilum Panicum crus-galli var. purpureum Panicum crus-galli var. pygmaeum Panicum crus-galli var. rohlenae Panicum crusgalli var. zelayense Echinochloa crus-galli f. rohlenae Echinochloa crus-galli f. submutica Echinochloa crus-galli f. vittata Echinochloa zelayensis Panicum crus-galli f. hispidum Panicum crus-galli f. exiguum Echinochloa crus-galli f. crus-galli Echinochloa macrocarpa var. mutica Echinochloa crusgalli var. breviseta Echinochloa crus-galli var. crus-galli Panicum crus-galli var. vulgare Panicum crus-galli var. angustifolia Echinochloa crus-galli var. aristata Echinochloa crus-galli var. crus-corvi Echinochloa crusgalli subsp. submutica Echinochloa crus-galli var. submutica Panicum crus-galli var. erectum Panicum crus-galli var. submuticum Echinochloa crus-galli subsp. erecta Echinochloa crus-galli

Lower taxons

Echinochloa crus-galli var. angustifolia Echinochloa crus-galli var. sieberiana