Epilobium ciliatum Raf.

American willowherb (en), Epilobe cilié (fr), Épilobe cilié (fr)

Species

Angiosperms > Myrtales > Onagraceae > Epilobium

Characteristics

Herbs perennial, erect, with compact leafy rosettes or rarely fleshy turions that leave brown basal scales. Stems (10-)25-90(-150) cm tall, well-branched or rarely simple, strigillose and glandular pubescent, sparsely pubescent or glabrescent below, with raised strigillose lines decurrent from margins of petioles. Leaves sessile above, lower ones with petioles 1-3 mm; cauline blade lanceolate to narrowly ovate, 2.5-6(-7) × 0.6-1.5(-2) cm, subglabrous except for strigillose margin and veins, base rounded or rarely subcordate, margin serrulate with 10-30 teeth per side, apex acute to subacuminate. Inflorescence and flowers erect. Sepals 2.4-3.5 mm, keeled. Petals pink or white, rarely rose-purple, 3.5-5(-7) mm. Stigma clavate to cylindric, entire. Capsules 4.5-7 cm, sparsely strigillose and glandular; pedicels 0.5-0.8(-1.4) cm. Seeds brown, 0.8-1.2 mm, with conspicuous longitudinal ridges of flattened, fused papillae, with chalazal collar 0.08-0.1 mm; coma dull white, readily detaching. Fl. Jul-Aug(-Sep), fr. Aug-Oct. 2n = 36.
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Erect herb 15-200 cm tall, increasing by leafy rosettes near the base of the thick, hollow stem; stems strigillose only along the elevated lines decurrent from the margins of the petioles. Lvs mostly opposite (the uppermost alternate), deep green, glabrous, (1)-2-9 × 0.7-3 cm, lanceolate, serrulate with 20-30 teeth each side; base rounded or obtuse, apex acuminate; petiole 1-2 mm long. Infl. nodding, strigillose and with glandular hairs. Fls erect. Ovary 0.8-2.5 cm long, hairy, sometimes densely so. Floral tube 0.5-1.4 mm deep, with a conspicuous ring of long hairs inside. Sepals keeled, 2-4 × 1-1.2 mm. Petals rose-purple, 3.5-5.5 × 2.2-3.2 mm. Filaments of the longer stamens 1.4-3 mm long. Style white, 1.5-3 mm long, glabrous; stigma clavate, 1.2-2 mm high. Capsules hairy, 4.5-8.5 cm long; fruiting pedicel 0.3-1 cm long. Seeds 0.8-1.2 × 0.3-0.45 mm, narrowly obovoid; abaxial surface with conspicuous lines of papillae; terminal end with an evident pellucid beak formed from the coalesced bases of the coma.
Perennating by persistent basal rosettes, without turions or long rhizomes; stems erect, mostly solitary, freely branched when well developed, to 1 or even 1.5 m, puberulent at least above, usually in decurrent lines, sometimes even sericeous, and often also glandular in the infl; lvs opposite, or the upper offset or alternate, lanceolate or lance-ovate, mostly 3–12 × 0.5–3.5 cm, serrulate, the teeth somewhat remote (usually 2–5 per cm of margin) or small and obscure; fls numerous; sep 2–6 mm, not projecting in bud; pet 2–6 mm, white (pink), notched; fr 4–10 cm, on a ± evident pedicel 2–15+ mm; seeds numerous, ca 1 mm, broadly short-beaked, longitudinally finely ribbed (evidently so at 20×); coma nearly white; 2n=36. Wet places, often in unstable habitats, variable and often somewhat weedy; Nf. and Lab. to Alas., s. to Va., w. N.C., Ind., Io., Calif., Tex., Mex., and C. Amer.; Chile and Arg. June–Aug. Ours is the widespread var. ciliatum. (E. adenocaulon; E. americanum; E. perplexans)
Erect perennial (occasionally annual) to c. 2 m high, with leafy shoots developing from the base. Stems with a rather close pubescence of irregularly curved hairs, restricted to raised lines decurrent from leaf bases, and often numerous glandular hairs, especially in the flowering region. Leaves usually opposite, alternate in upper part of the stem, lanceolate, 2.5–8 cm long, with numerous very shallow teeth, the apex tapering or acuminate, the base broad and rounded. Flowers with a conspicuous ring of long hairs inside hypanthium. Petals 3.5–5.5 mm long, purplish pink or white. Capsule 4.5–8.5 cm long, with a close partly glandular pubescence; pedicel 0.3–1 cm long. Seeds very narrowly obovoid, 0.9–1.2 mm long, the surface with obvious lines of laterally flattened papillae.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination autogamy
Spread anemochory
Mature width (meter) 0.1 - 0.5
Mature height (meter) 0.25 - 1.2
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Light 4-9
Soil humidity 5-7
Soil texture 3-4
Soil acidity 2-8
Soil nutriment -
Hardiness (USDA) 7-10

Usage

Uses medicinal
Edible -
Therapeutic use Orthopedic Aid (root), Antidiarrheal (root), Analgesic (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings, divisions or seedlings.
Mode cuttings divisions seedlings
Germination duration (days) 14 - 30
Germination temperacture (C°) 10 - 15
Germination luminosity light
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Epilobium ciliatum habit picture by Sabina Hartmann (cc-by-sa)
Epilobium ciliatum habit picture by Lisa J (cc-by-sa)
Epilobium ciliatum habit picture by David Hocken (cc-by-sa)

Leaf

Epilobium ciliatum leaf picture by Konrad Preis (cc-by-sa)
Epilobium ciliatum leaf picture by Stefan Steinbach (cc-by-sa)
Epilobium ciliatum leaf picture by Dries de Moor (cc-by-sa)

Flower

Epilobium ciliatum flower picture by soulhuntress (cc-by-sa)
Epilobium ciliatum flower picture by Uwe Dr. Heitz (cc-by-sa)
Epilobium ciliatum flower picture by William Sixsmith (cc-by-sa)

Fruit

Epilobium ciliatum fruit picture by Christian Henkel (cc-by-sa)
Epilobium ciliatum fruit picture by Anke Vieth (cc-by-sa)
Epilobium ciliatum fruit picture by Jenn (cc-by-sa)

Distribution

Epilobium ciliatum world distribution map, present in Argentina, Australia, Austria, Belgium, Belarus, Canada, Switzerland, Chile, China, Czech Republic, Germany, Estonia, Finland, Falkland Islands (Malvinas), France, Greece, Guatemala, Hungary, Italy, Liechtenstein, Lithuania, Luxembourg, Latvia, Mexico, Netherlands, Norway, New Zealand, Poland, Russian Federation, Slovakia, Sweden, Uruguay, and United States of America

Conservation status

Epilobium ciliatum threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:611725-1
WFO ID wfo-0000668926
COL ID 6G4VY
BDTFX ID 24079
INPN ID 96149
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Epilobium adenocaulon f. umbrosum Epilobium chilense f. verticillatum Epilobium alaskae Epilobium albiflorum Epilobium constrictum Epilobium delicatum Epilobium deminutum Epilobium ecomosum Epilobium novomexicanum Epilobium ovatum Epilobium palmeri Epilobium rubescens Epilobium ursinum Epilobium valdiviense Chamaenerion domini Epilobium aconcaguinum Epilobium adenocladon Epilobium affine Epilobium americanum Epilobium brevistylum Epilobium caesiovirens Epilobium chilense Epilobium concinnum Epilobium cunninghamii Epilobium domini Epilobium fendleri Epilobium griseum Epilobium leiophyton Epilobium longipes Epilobium macdougalii Epilobium magellanicum Epilobium praecox Epilobium pseudolineare Epilobium santa-cruzense Epilobium perplexans Epilobium chilense f. latifolium Epilobium chilense f. macrum Epilobium californicum Epilobium holosericeum Epilobium hookerianum Epilobium parishii Epilobium valdiviense f. elatius Epilobium valdiviense f. simplex Epilobium adenocaulon subsp. americanum Epilobium adenocaulon var. ecomosum Epilobium adenocaulon var. holosericeum Epilobium adenocaulon var. parishii Epilobium adenocaulon var. pseudocoloratum Epilobium adenocaulon subsp. rubescens Epilobium brevistylum var. ursinum Epilobium californicum var. holosericeum Epilobium californicum var. parishii Epilobium chilense var. latifolia Epilobium chilense var. latifolium Epilobium chilense var. macra Epilobium chilense var. macrum Epilobium chilense var. valdiviense Epilobium ciliatum var. ecomosum Epilobium ciliense var. valdiviense Epilobium glandulosum var. asiaticum Epilobium glandulosum var. ecomosum Epilobium pedicellare var. latifolium Epilobium santa-cruzense var. cruzense Epilobium valdiviense var. alboffii Epilobium watsonii var. parishii Epilobium adenocaulon var. americanum Epilobium glandulosum var. kurilense Epilobium adenocaulon f. albiflorum Epilobium adenocaulon f. apricum Epilobium ciliatum subsp. ciliatum Epilobium chilense var. verticillata Epilobium argentium Epilobium ciliatum

Lower taxons

Epilobium ciliatum subsp. watsonii