Epilobium L.

Willowherb (en), Épilobe (fr)

Genus

Angiosperms > Myrtales > Onagraceae

Characteristics

Erect or creeping herbs, usually perennial, very rarely annual, sometimes woody near base, overwintering and multiplying by leafy rosettes, subterranean or surface runners and stolons, turions or gemmae. Lvs opposite, at least below infl., often lower opposite and upper alternate, occasionally all alternate, rarely verticillate, simple. Fls solitary in lf axils, but sometimes forming a ± definite, terminal, racemose infl., usually actinomorphic, occasionally somewhat zygomorphic; pollen and stigmas ripening together, occasionally markedly protogynous or protandrous. Buds erect to pendulous. Floral tube present or 0, cylindric. Sepals 4, caducous. Petals 4, nearly always white or cream to pink, rose or purple, usually emarginate, sometimes entire, not clawed. Stamens 8, the outer whorl longer; anthers basifixed or dorsifixed. Stigma capitate, clavate, or deeply 4-lobed. Capsule 4-celled, loculicidal, slender. Seeds usually numerous, ± brown, nearly always with terminal, caducous or persistent coma of silky hairs.
More
Herbs perennial [or annual, sometimes suffrutescent], with leafy rosettes, stolons, soboles (shoots), or turions (subterranean globose buds with fleshy scales). Stems glabrous to pubescent, often with lines of hairs decurrent from margins of petioles. Leaves opposite, becoming alternate and bractlike in inflorescence; petiolate or sessile; stipules absent; bracteoles absent. Inflorescences simple or branched racemes, panicles, spikes, or corymbs. Flowers 4-merous, often protandrous, with floral tube, producing nectar at base of style. Petals pink to rose-purple or white [or rarely cream-colored or orange-red], obcordate or obtrullate, notched at apex. Stamens 8, in two unequal whorls; pollen yellow, shed in tetrads. Style erect; stigma entire or 4-lobed. Fruit an elongate, slender capsule, 4-loculed, loculidical. Seeds many [or rarely only 4], generally with terminal coma of silky hairs [or coma rarely lacking]. 2n = [18, 20, 24, 26, 28, 30, 32,] 36, [38, 60].
Perennial herbs, often flowering in the first year, occasionally somewhat woody near the base. Leaves (in Mal.) opposite below, spirally arranged above. Flowers in the axils of the often greatly reduced upper leaves. Floral tube short (in Mal.) or essentially absent. Sepals 4, erect. Petals 4, white, pink, or purple, emarginate (in Mal.). Stamens 8, in 2 whorls, the epipetalous ones shorter. Stigma (in Mal.) clavate or rarely globose, usually surrounded by the shedding anthers at maturity. Ovary 4-locular, the ovules very numerous. Fruit a long, slender, loculicidal capsule. Seeds very numerous, small, with a chalazal plume of trichomes (coma).
Fls 4-merous; hypanthium usually prolonged a little beyond the summit of the elongate, slender ovary and simulating a cal-tube; sep ovate to lanceolate, deciduous from the fr; pet white to pink or purple; stamens 8, often unequal; style short; stigma clavate, ovoid, or 4-cleft; capsule linear, elongate, 4-valved; seeds numerous, small, bearing a terminal coma of hairs; annual or usually perennial herbs, rarely subshrubs, with alternate or opposite, linear to ovate lvs; fls many to few in elongate or short terminal racemes, or even solitary. 200, N. and S. Temp. and Arctic.
Perennial or rarely annual herbs, sometimes woody at base. Leaves opposite or alternate. Flowers solitary in upper leaf axils, usually actinomorphic; without bracteoles. Hypanthium short but evident or (not in Australia) absent or to 4 cm long. Sepals 4, deciduous. Petals 4, usually white, pink or purplish pink, or (not in Australia) yellow or red. Stamens 8. Ovary 4-locular. Fruit a slender, elongate capsule. Seeds numerous, occasionally (not in Australia) few, usually with a coma of long, fine hairs.
Fls us. solitary, regular, 4-merous; perig. with very short calyx-tube; petals us. 2-lobed; stamens 8, in 2 series; ovary 4-celled, ovules ∞; stigma capitate to 4-lobed. Fr. a long narrow capsule with loculicidal dehiscence; seeds mostly papillose, with a chalazal brush of hairs (coma). Herbs or subshrubs with at least lower lvs opp. Some 200 spp., mainly of temperate regions.
Plants herbaceous; stems creeping and rooting in basal part, then erect or ascending, sts slightly woody at very base. Fls us. few, confined to axils of uppermost lvs; stigma us. clavate.
Stock ± woody; stems elongate, erect, herbaceous, many-fld towards tips; lvs rather large, of oblong or lanceolate order. Stigma us. clavate.
Stems slender, herbaceous, prostrate, rooting at nodes; fls in axils of intermediate lvs; mature peduncles much > lvs; stigma clavate.
Flowers in a terminal inflorescence (in F.Z. area) or borne in the axils of the leaves.
Petals 4, rose-purple or white (yellow in 1 N. American species), emarginate.
Ovary 4-locular; ovules pluriseriate in each locule, the seeds free.
Perennial herbs, often flowering the first year, rarely annual.
Pollen shed in tetrads, singly in a few non-African species.
Leaves opposite, usually alternate above, toothed or entire.
Seeds with a prominent tuft of hairs, the coma.
Stigma clavate, globose, or deeply 4-lobed.
Floral tube present (in F.Z. area) or nil.
Dehiscence of capsule loculicidal.
Stamens 8; anthers versatile.
Sepals 4, not persistent.
Bracteoles lacking.
Stipules absent.
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system creeping-root
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Characteristic of open, disturbed habitats or grassland or the alpine zone, not normally found in primary forest.Dispersal takes easily place by wind as the light seeds are provided with a silky coma.It is still rather surprising that, whereas open habitats are very common in the Malesian mountains, with their numerous volcanoes and alpine habitats, the number of localities is so very restricted. It must probably be considered that though dispersal may be easy, the coma only acts efficiently in dry air and this is in the tropics with their frequent rain and cloud formation and nightly fogs during most of the year a rather rare climatic situation. In this respect the west-monsoon, blowing from Asia southeastwards (Nov.-March) is most unsuitable, as these winds are laden with moisture. The east-monsoon, blowing from Australia northeastwards (May-Sept.) is in this respect definitely more favourable as this is a dry wind. This wind regime is a consequence of the situation of the Asian and Australian continents.Local dispersal by water might also occur, as the lowest localities in New Guinea are all along stream-sides, but this may also be due to the downward air-stream over rushing water.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 4-10

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 22 - 45
Germination temperacture (C°) 13 - 17
Germination luminosity light
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Epilobium unspecified picture

Distribution

Epilobium world distribution map, present in Australia, China, Indonesia, Sri Lanka, New Zealand, Philippines, Papua New Guinea, Timor-Leste, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:30000954-2
WFO ID wfo-4000013609
COL ID 4D6C
BDTFX ID 86380
INPN ID 192232
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Zauschneria Chamaenerion Boisduvalia Chamerion Epilobium

Lower taxons

Epilobium campestre Epilobium subdentatum Epilobium adenocaulon Epilobium aitchisonii Epilobium anglicum Epilobium angustum Epilobium astonii Epilobium atlanticum Epilobium billardierianum Epilobium x borbasianum Epilobium brunnescens Epilobium capense Epilobium colchicum Epilobium x confine Epilobium conspersum Epilobium curtisiae Epilobium cylindricum Epilobium davuricum Epilobium x decipiens Epilobium densiflorum Epilobium densifolium Epilobium denticulatum Epilobium duriaei Epilobium x finitimum Epilobium gemmascens Epilobium x gemmiferum Epilobium glaberrimum Epilobium gunnianum Epilobium hallianum Epilobium x haussknechtianum Epilobium hectori Epilobium hirsutum Epilobium x huteri Epilobium leptocarpum Epilobium leptophyllum Epilobium limosum Epilobium luteum Epilobium marshallianum Epilobium minutum Epilobium mirabile Epilobium montaniforme Epilobium neogradiense Epilobium nerterioides Epilobium nevadense Epilobium nivale Epilobium nivium Epilobium x novae-civitalis Epilobium nummulariifolium Epilobium nutans Epilobium obcordatum Epilobium oreganum Epilobium palatinum Epilobium pallidiflorum Epilobium pallidum Epilobium pedunculare Epilobium pengii Epilobium x percollinum Epilobium pernitens Epilobium perpusillum Epilobium pseudotrigonum Epilobium psilotum Epilobium puberulum Epilobium rechingeri Epilobium x reedii Epilobium rigidum Epilobium x rivulare Epilobium roseum Epilobium rostratum Epilobium rotundifolium Epilobium royleanum Epilobium salignum Epilobium saximontanum Epilobium septentrionale Epilobium x sericeum Epilobium sikkimense Epilobium stevenii Epilobium strictum Epilobium suffruticosum Epilobium taiwanianum Epilobium tonkinense Epilobium trichophyllum Epilobium x uechtritzianum Epilobium verticillatum Epilobium wallichianum Epilobium wilsonii Epilobium chionanthum Epilobium waterfallii Epilobium wattianum Epilobium williamsii Epilobium willisii Epilobium abortivum Epilobium aggregatum Epilobium alsinoides Epilobium amurense Epilobium australe Epilobium blinii Epilobium x boissieri Epilobium brachiatum Epilobium brevifolium Epilobium brevipes Epilobium x brevipilum Epilobium brevisquamatum Epilobium x brunnatum Epilobium x celakovskyanum Epilobium x chateri Epilobium chitralense Epilobium chlorifolium Epilobium clarkeanum Epilobium clavatum Epilobium cleistogamum Epilobium confertifolium Epilobium confusum Epilobium conjungens Epilobium detznerianum Epilobium x erroneum Epilobium x fallacinum Epilobium fangii Epilobium fastigiato-ramosum Epilobium fauriei Epilobium x floridulum Epilobium forbesii Epilobium x fossicola Epilobium fragile Epilobium freynii Epilobium frigidum Epilobium fugitivum Epilobium gerstlaueri Epilobium glabellum Epilobium glaciale Epilobium x glanduligerum Epilobium gouldii Epilobium x haynaldianum Epilobium hectori-leveilleanum Epilobium heterocaule Epilobium hirtigerum Epilobium hohuanense Epilobium hooglandii Epilobium hornemannii Epilobium howellii Epilobium indicum Epilobium insulare Epilobium x interjectum Epilobium x intersitum Epilobium x jinshaense Epilobium karsteniae Epilobium x keredjense Epilobium kermodei Epilobium keysseri Epilobium kingdonii Epilobium x kitcheneri Epilobium komarovianum Epilobium komarovii Epilobium korshinskyi Epilobium krausei Epilobium ladakhianum Epilobium x lamotteanum Epilobium x langeanum Epilobium x laschianum Epilobium latifolium Epilobium laxum Epilobium leiophyllum Epilobium lipschitzii Epilobium macropus Epilobium madrense Epilobium margaretae Epilobium matthewsii Epilobium maysillesii Epilobium melanocaulon Epilobium x mentiens Epilobium mexicanum Epilobium microphyllum Epilobium minutiflorum Epilobium nankotaizanense Epilobium x nutantiflorum Epilobium x obscurescens Epilobium oregonense Epilobium pannosum Epilobium parviflorum Epilobium platystigmatosum Epilobium ponticum Epilobium prionophylloides Epilobium prostratum Epilobium purpuratum Epilobium purpureum Epilobium pycnostachyum Epilobium pyrricholophum Epilobium rhynchospermum Epilobium x rivulicola Epilobium sarmentaceum Epilobium schmidtianum Epilobium x schulseanum Epilobium semiamplexicaule Epilobium shiroumense Epilobium x simulatum Epilobium sinense Epilobium siskiyouense Epilobium spitianum Epilobium stereophyllum Epilobium subalgidum Epilobium subcoriaceum Epilobium tasmanicum Epilobium thermophilum Epilobium x thuringiacum Epilobium tianschanicum Epilobium tibetanum Epilobium turkestanicum Epilobium x udicola Epilobium vernonicum Epilobium winkleri Epilobium x wisconsinense Epilobium sericeum Epilobium x treleasianum Epilobium x krausei Epilobium x aschersonianum Epilobium x vicinum Epilobium amphibolum Epilobium pedicellare Epilobium lactiflorum Epilobium lanceolatum Epilobium hohuanense Epilobium algidum Epilobium anatolicum Epilobium x confusilobum Epilobium x cornubiense Epilobium crassum Epilobium dacicum Epilobium dasycarpum Epilobium glaucum Epilobium goerzii Epilobium gracilipes Epilobium grenieri Epilobium griffithianum Epilobium persicinum Epilobium porphyrium Epilobium stracheyanum Epilobium pubens Epilobium x argillaceum Epilobium arvernense Epilobium barbeyanum Epilobium pseudorubescens Epilobium speciosum Epilobium staintonii Epilobium decipiens Epilobium facchini Epilobium x ludmilae Epilobium petraeum Epilobium pictum Epilobium brachycarpum Epilobium coloratum Epilobium fleischeri Epilobium canum Epilobium obscurum Epilobium anagallidifolium Epilobium collinum Epilobium alpestre Epilobium palustre Epilobium alsinifolium Epilobium ciliatum Epilobium dodonaei Epilobium tetragonum Epilobium montanum Epilobium angustifolium