Herbs, annual, with taproot. Stems prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). Leaves opposite; stipules distinct, linear-subulate, often irregularly 2-or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. Cyathia solitary at distal nodes; peduncle 0.1–0.6(–2) mm. Involucre obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. Staminate flowers 2–5. Pistillate flowers: ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. Capsules ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. Seeds white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate.