Herbs, annual or perennial, often robust, with slender to moderately thickened taproot. Stems erect to ascending, 30–100 cm, pilose to tomentose. Leaves opposite; stipules distinct when young, connate into deltate scale when older, erose to laciniate, with dark glands at margin or base, 0.5–1.3 mm, pilose; petiole 0.5–2 mm, glabrescent or pilose; blade ovate or oblong, sometimes slightly falcate, 8–46 × 3–21 mm, base asymmetric, obtuse to hemicordate, margins serrulate, apex obtuse or acute, surfaces often with red spot in center, pilose to sericeous; palmately 3–5(–7)-veined from base. Cyathia in dense, usually terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia; peduncle 0.8–2.3 mm. Involucre obconic, 0.9–1.3 × 0.8–1.2 mm, pilose to sericeous; glands 4, yellow or pink, circular or oblong, 0.1–0.3 mm diam.; appendages white or pink, oblong, flabellate, or suborbiculate, 0.1–0.6 × 0.2–1 mm, entire or crenate. Staminate flowers 15–25. Pistillate flowers: ovary densely pilose to sericeous with yellowish hairs; styles 0.6–0.9 mm, 2-fid 1/2 length, filiform. Capsules subglobose to broadly ovoid, 1.7–2 mm diam., pilose to sericeous with yellowish hairs; columella 1.5–1.7 mm. Seeds dark reddish brown to almost black, plumply ovoid, 4-angled in cross section, 1.2–1.3 × 0.7–0.8 mm, with 2 inconspicuous rows of 3–5 shallow depressions separated by low ridges.
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