Herbs, perennial, with thick, globose to elongated tubers, 2–8 cm. Stems erect to ascending, branched, 10–45(–60) cm, glabrous, puberulent, or densely hirsute to setose, often with 2-layered indumentum of long hairs intermixed with short hairs. Leaves usually opposite, occasionally whorled distally, or rarely with 1–2 alternate leaves; stipules 0.1–0.2 mm; petiole 0–18 mm, hirsute, sericeous, or strigose; blade linear to ovate or almost orbiculate, 6–54 × 2–19 mm, base rounded to attenuate, margins entire, occasionally ciliate with stiff recurved hairs, apex acute to obtuse, surfaces usually hirsute, sericeous, or strigose, occasionally glabrous adaxially; venation conspicuous. Cyathia in weakly-defined terminal dichasia; peduncle 1.4–5.8 mm, glabrous. Involucre obconic to campanulate, 1.1–1.4 × 0.5–1.5 mm, glabrous or strigillose; glands 4–5, greenish, oblong, 0.2 × 0.4–0.5 mm; appendages usually yellowish or green, rarely dark purple, ovate, flabellate, semiorbiculate, or oblong, 0.3–0.9 × 0.4–1.1 mm, usually entire. Staminate flowers 10–15. Pistillate flowers: ovary glabrous, sericeous, or strigillose; styles 0.4–0.6 mm, 2-fid 1/2 length. Capsules oblate, 2.3–3 × 3.1–4.2 mm, glabrous, sericeous, or strigillose; columella 1.6–2.1 mm. Seeds black to light brown, broadly ovoid to subglobose, rounded in cross section, 1.5–2.3 × 1.4–1.8 mm, smooth or with low rounded tubercles; caruncle absent.
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