Herbs, annual, with slender taproot. Stems erect to ascending, 10–55 cm, glabrous. Leaves opposite; stipules distinct, narrowly triangular and often divided into 3–5 subulate to filiform segments, 0.8–1.9 mm, glabrous; petiole 1.2–3.1 mm, glabrous; blade oblong, elliptic, or ovate, 12–43 × 3–14 mm, base subsymmetric to asymmetric, rounded, attenuate, or with one side hemicordate and other side rounded, margins usually serrulate, rarely entire on some leaves or portion of blade, apex obtuse to acute, surfaces often with red blotch in center, glabrous; pinnately veined, often only midvein conspicuous. Cyathia solitary at distal nodes; peduncle 1.4–2.4 mm. Involucre oblong, campanulate, or funnel-shaped, 1.5–2.6 × 1.1–2.3 mm, glabrous; glands 4, green to yellowish or tinged with red, subcircular to oblong, 0.4–0.7 × 0.5–1 mm; appendages absent or white, often forming narrow rim or oblong to ovate, 0–0.8 × 1.2 mm, distal margin entire. Staminate flowers 50–60. Pistillate flowers: ovary glabrous; styles 0.5–0.6 mm, 2-fid 1/2 length. Capsules oblate, 2.4–3.2 × 3–3.5 mm, glabrous; columella 2.3–2.5 mm. Seeds light brown, ovoid, 4-angled in cross section, abaxial keel well developed, 1.9–2.3 × 1.3–1.4 mm, finely dimpled and papillate.
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