Herbs, perennial, with thick rootstock. Stems slender, erect or ascending, sometimes sinuous, densely branched near base, 10–50 cm, moderately to densely puberulent to lanulose. Leaves: petiole 0–1 mm; blade usually lanceolate or elliptic-lanceolate, sometimes slightly oblanceolate, 8–30 × 6–14 mm, base usually acute, occasionally short-attenuate, rarely obtuse, margins entire, apex usually acute, occasionally obtuse, acuminate, or cuspidate, surfaces sparsely to moderately puberulent to lanulose; venation pinnate, sometimes obscure, midvein prominent. Cyathial arrangement: terminal pleiochasial branches 3–5, each 1–2 times 2-branched; pleiochasial bracts broadly ovate to subcordate, usually similar in size to, occasionally wider than, distal leaves; dichasial bracts distinct, broadly ovate to almost reniform, base obtuse, margins entire, apex obtuse, acuminate to cuspidate; axillary cymose branches 0–5. Cyathia: peduncle 0.3–0.8 mm. Involucre campanulate to broadly turbinate, 2.2–3 × 2–2.5 mm, puberulent to lanulose; glands 4, semicircular, trapezoidal, or elliptic-truncate, 0.8–1.5 × 1–2.2 mm, margins strongly crenate or dentate; horns usually absent, if present then straight, 0.1–0.2 mm, generally equaling teeth on gland margin. Staminate flowers 12–20. Pistillate flowers: ovary usually puberulent, occasionally lanulose; styles 1–1.2 mm, 2-fid. Capsules oblong-ovoid, 3.5–4 × 3–4 mm, 3-lobed; cocci rounded, smooth, usually puberulent, occasionally lanulose; columella 2.5–3 mm. Seeds gray to whitish, oblong cylindric, 2–3 × 1.5–1.8 mm, irregularly shallowly pitted to almost smooth; caruncle conic, 0.6 × 0.6 mm.