Herbaceous or suffruticose, lactiferous vines; stems terete, pubescent, the hairs of 2 types, short brown puberulent hairs overtopped by brown, septate, pilose hairs to 5 mm long of variable density but rarely lacking. Leaves petiolate, membranaceous to coriaceous, sparsely to densely pubescent on both surfaces, especially along the major veins, usually basally cordate, the sinuses narrow, the lobes frequently imbricate, usually with a cluster of digitiform glands at the juncture of the petiole and blade. Inflorescences interaxillary, at first umbelliform, becoming racemose, brown puberulent throughout with short hairs usually overtopped by longer ones. Flowers pedicellate; calyx deeply 5-lobed, abaxially puberulent or puberulent and pilose, adaxially glabrous or pubescent, 1 or 2 digitiform glands present in each sinus, lobes ovate to lanceolate or subulate; corolla subrotate, greenish, yellowish, or white, the tube short, usually not ex-ceeding 2 mm, lobes ovate to suborbicular, canaliculate or plane, abaxially short-strigose to glabrate, glabrous along those parts of the margins which are im-bricate in bud, adaxially pubescent, the hairs usually flattened, septate, and whitish, the surfaces frequently papillate beneath the pubescence; stamens connate to the top of the corona, above the corona merely connivent, corona carnose, lobed, angular, or entire, longitudinally striate-sulcate, fimbriate, rarely smooth, basally adnate to the corolla and for its full length to the staminal fila-ments, apically abutting the underside of the peltate stigma head and held in position by the inflexed terminal membranes which wholly or partially cover the top of the stigma head; stigma head broadly 5-angled, plane or slightly depressed in the center, the styles short, the ovaries obliquely ovoid to ellipsoid, glabrous to densely pubescent, only one developing in fruit; pollinia inserted horizontally or nearly so, subreniform, the fertile portion + lunate, the dorsal margins pellucid, sterile. Follicles obliquely ovoid to ellipsoid, glabrous or minutely pubescent, smooth or striate; seeds compressed ovoid, the marginal wings coarsely and irregularly toothed in the lower half.
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