Secondary stems irregularly pinnate; paraphyllia absent; pseudoparaphyllia subfoliose to occasionally filamentous. Stem leaves loosely appressed to imbricate when dry, erect-spreading to squarrose when moist, lanceolate, ovate-lanceolate or broadly so, ovate, ovate-acuminate, oval, or deltoid; margins recurved to mid leaf or beyond, entire or serrulate distally; apex acute to short-acuminate; costa single or double, ending before apex; alar region extending up margins to 1/3 leaf length; medial laminal cells elliptic to linear. Branch leaves similar, smaller. Sexual condition autoicous; perichaetial leaf apex straight. Seta 0.4-3.9 mm. Capsule erect, immersed to exserted, cylindric; stomata absent; operculum conic to erect or obliquely rostrate; exostome teeth free to base or rarely connate proximally, linear-lanceolate, smooth, granulose, or minutely papillose; endostome rudimentary to absent. Calyptra hairy. Spores 15-35 µm, green.
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