Plants small to large, light green to yellow-green. Stems creeping, irregularly pinnate, branches arcuate to erect, short to elongate, simple to many-branched, incurved or rigid when dry, in clusters of 1-5 on either side of vegetative growth interval; paraphyllia present or absent; pseudoparaphyllia subfoliose to filamentous, entire; axillary hairs 6-8 per axil, proximal cells 1 or 2, short, walls light brown, distal cells 2-4, elongate, hyaline. Leaves spirally inserted, loosely appressed to imbricate when dry, erect-spreading to squarrose when moist, rapidly spreading when moistened, usually ovate-lanceolate; margins entire or serrate distally; apex obtuse, acute, or acuminate; costa single, ending before apex, or double, short; alar cells quadrate to transversely elongate; laminal cell walls thick; medial cells isodiametric to linear, smooth. Specialized asexual reproduction absent. Sexual condition autoicous or dioicous; perigonia lateral, gemmiform; perichaetia lateral, larger, inner leaves following fertilization, paraphyses elongating. Seta single. Capsule immersed to exserted, symmetric; stomata absent, or basal, sunken; annulus absent; operculum conic-rostrate; peristome usually double, pale; exostome teeth 16; endostome rudimentary or absent to well developed, cilia absent. Calyptra cucullate, hairy or naked. Spores spheric, minutely papillose.
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