Trees with simple long-petioled undivided or 3–5-lobed leaves. Flowers very small and numerous in corymbose ebracteate cymes borne chiefly in the axils of the upper leaves. Flowers polygamous, mostly male. Perianth 4–8-partite; lobes in a single whorl, imbricate, equal or unequal, 2 larger than the rest and accrescent in hermaphrodite flowers. Stamens much exserted, 2–7 but usually 4, inserted at the base of the perianth; anthers dehiscing laterally, the valves opening upwards. Staminodes ± clavate, as many as and alternate with the stamens, or more numerous. Style straight; stigma obliquely expanded. Fruit with 2 terminal wings formed by accrescent perianth-segments. Cotyledons spirally twisted.
Trees, monoecious. Leaves simple, sometimes palmately lobed. Inflorescence an axillary panicle, often corymbose. Flowers bisexual or unisexual. Perianth lobes 4–8, usually 7, minute. Bisexual flowers: stamens 4–7, exserted, usually alternating with as many staminodes; anther filaments usually glandular; style terete; stigma capitate. Male flowers: stamens as in bisexual flowers; pistillode sometimes present. Female flowers: gynoecium as in bisexual flowers; staminodes usually present. Fruit nut-like, usually ± ribbed longitudinally, with 2 apical, spathulate wings.
Perianth segments in one 4–8-merous imbricate whorl, equal or unequal with 2 segments larger and accrescent in hermaphrodite flowers.
Stamens (2)4(7), exserted, inserted at base of perianth; anthers dehiscing laterally by upward-opening valves.
Fruit a globose ovoid or ellipsoid drupe with 2 long apical wings formed by accrescent perianth segments.
Flowers small, numerous, hermaphrodite and unisexual on the same individual, predominantly male.
Inflorescences of corymbose, ebracteate cymes, mainly in upper leaf axils.
Staminodes as many as and alternating with the stamens, or more numerous.
Leaves simple, undivided or 3–5-lobed, long-petioled.
Ovary and style reduced or absent in male flowers.
Deciduous trees, rarely shrubs.
Cotyledons spirally twisted.